Bakgrunn og aktiviteter



BI2044 Etologi - kursansvarlig

BI3040 Atferdsøkologi - kursansvarlig

BI3052 Study Design in Biology - kursansvarlig



Evolusjon av samarbeide og konflikt, foreldreomsorg, konflikt mellom foreldre og avkom, og samarbeide hekking på fugler.

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PUBLIKASJONER (.pdf-filer tilgjengelig ved henvendelse)

  • Wright, J. & McDonald, P G. (2016) Kin selected helping decisions of the bell miner. In: Cooperative Breeding: Studies of Ecology, Evolution, and Behavior. (Ed by Koenig, W. & Dickinson, J.) Cambridge University Press, Cambridge.
  • Westneat, D.F., Wright, J. & Dingemanse, N.J. (2015) The biology hidden within residual phenotypic variance. Biol. Rev. 90, 729-743.
  • Botero, C.A., Weissing, F.J., Wright, J. & Rubenstein, D.R. (2015) Evolutionary tipping points in the capacity to adapt to environmental change. PNAS 112, 184-189.
  • Pélabon, C., Albertsen, E., Falahati-Anbaran, M., Wright, J. & Armbruster, W.S. (2015) Does multiple paternity affect seed size in angiosperms? An experimental test in Delachampia scandensJ. Evol. Biol. 28, 1719-1733.
  • Ratikainen, I.I. & Wright, J. (2013) Adaptive management of body mass in Siberian jays. Anim. Behav. 85, 427-434.
  • Westneat, D.F., Schofield, M. & Wright, J. (2013) Parental behavior exhibits between-individual variance, plasticity and heterogeneous residual variance. Behav. Ecol. 24, 598-604.
  • Mutzel, A., Blom, M.P.K., Spagopoulou, F., Wright, J., Dingemanse, N.J. & Kempenaers, B. (2013) Temporal trade-offs between nestling provisioning and defence against nest predators in blue tits. Anim. Behav. 85, 1459-1469.
  • Dingemanse, N.J., Barber, I., Wright, J. & Brommer, J. (2012) Quantitative genetics of behavioural reaction norms: genetic correlations between personality and behavioural plasticity vary across stickleback populations. J. Evol. Biol. 25, 485-496.
  • Mathot K.J., Wright, J, Kempenaers, B., & Dingemanse, N.J. (2012) Adaptive strategies for managing uncertainty may explain personality-related differences in behavioural plasticity. Oikos 121, 1009-1020.
  • Ratikainen, I.I., Henøen, L.R., Wright, J. & Kazem, A.J.N. (2012) Private and public information by foraging groups of wild Siberian jays. Anim. Behav. 83, 1243-1251.
  •  Briga, M., Pen, I. & Wright, J.  (2012) Care for kin: within-group relatedness and allomaternal care are positively correlated and conserved throughout the mammalian phylogeny. Biol. Lett. 8, 533-536.
  •  Burger, C., Belskii, E., Eeva, T., Laaksonen, T., Mägi, M., Mänd, R., Qvarnström, A., Slagsvold, T., Veen, T., Visser, M.E., Wiebe, K.L., Wiley, C., Wright J., and Both, C. (2012) Climate change, breeding date and nestling diet: how temperature differentially affects seasonal changes in pied flycatcher diet depending on habitat variation. J. Anim. Ecol. 81, 926-936.
  • Wright, J. (2011) Honest begging: signals of need, quality and/or hunger. Behav. Ecol. 22, 920-921.
  • McDonald, P. G. & Wright, J. (2011) Bell miner provisioning calls are more similar among relatives and are used by helpers at the nest to bias their effort towards kin. Proc. Roy. Soc. Lond. B. 278, 3403-3411.
  • Réale, D., Dingemanse, N.J., Kazem, A.J.N. & Wright, J. (2010) Evolutionary and ecological approaches to the study of personality. Phil. Trans. Roy. Soc. Lond. B., 365, 3937-3946.
  • Ratikainen, I.I., Wright, J. & Kazem, A.J.N. (2010) Social class influences degree of variance-sensitivity in wild Siberian jays. Behav. Ecol. 21, 1067-1072.
  • Wright, J. & Radford, A.N. (2010) Risk-sensitive green woodhoopoes: a new explanation for sex-specific differences in preferred foraging techniques. Ethology 116, 941-950.
  • Rollins, L.A., Holleley, C.E., Wright, J., Russell, A.F., & Griffith, S.C. (2010) Isolation and characterization of polymorphic tetranucleotide microsatellite loci in the Apostlebird (Struthidea cinerea). Cons. Gen. Resour., 2, 229-231.
  • Wright, J., Karasov, W.H., Kazem, A.J.N., Goncalves, I.B. & McSwan, E. (2010) Begging and digestive responses to differences in long-term and short-term need in nestling pied flycatchers. Anim. Behav. 80, 517-525.
  • Dingemanse, N.J., Dochtermann, N. & Wright, J. (2010) A method for exploring the structure of behavioural syndromes to allow formal comparison within and between datasets. Anim. Behav. 79, 439-450.
  • Dingemanse, N.J., Kazem, A.J.N., Réale, D. & Wright, J. (2010) Behavioural reaction norms: animal personality meets individual plasticity. TREE 25, 81-89.
  • McDonald, P.G., Ewen, J.G. & Wright, J. (2010) Brood sex ratio does not affect helper effort in a cooperative bird, despite extreme sex-biased dispersal. Anim. Behav., 79, 243-250.
  • Wright, J., McDonald, P.G., te Marvelde, L., Kazem, A.J.N. & Bishop, C.M. (2010) Helping effort increases with relatedness in bell miners, but ‘unrelated’ helpers of both sexes still provide substantial care. Proc. Roy. Soc. Lond. B. 277, 437-445.
  • Lykkja, O.N., Solberg, E.J., Herfindal, I., Wright, J., Rolandsen, C.M. & Hanssen, M.G. (2009) The effects of human activity on summer habitat use by moose. Alces 45, 109-124.
  • Griesser, M., Barnaby, J., Schneider, N.A., Figenschau, N., Wright, J., Griffith, S.C., Kazem, A..J.N. & Russell, A.F. (2009) Winter ranging behaviour and the organization of cooperative breeding groups in the apostlebird. Ethology 115, 888-896.
  • Russell, A.F. & Wright, J. (2009) Avian mobbing: by-product mutualism not reciprocal altruism. TREE 24. 3-5.
  • van de Pol, M. & Wright, J. (2009) A simple method for distinguishing within- versus between-subjects effects using mixed models. Anim. Behav. 77, 753-758.
  • te Marvelde, L., McDonald, P.G., Kazem, A.J.N. & Wright, J. (2009) Do helpers really help? Biomass of different food types delivered and nestling growth rates in the cooperatively breeding bell miner. Anim. Behav. 77, 727-735.
  • McDonald, P.G., Kazem, A.J.N. & Wright, J. (2009) Cooperative provisioning dynamics: fathers and unrelated helpers show similar responses to manipulations of begging. Anim. Behav. 77, 369-376.
  • Dall, S.X.R. & Wright, J. (2009) Rich pickings near large communal roosts favor 'gang' foraging by juvenile common ravens, Corvus corax. PLOS ONE 4(2), e4530. doi:10.1371/journal.pone.0004530.
  • Dingemanse, N.J., Van der Plas, F., Wright, J., Réale, D., Schrama, M., Roff, D.A., Van der Zee E., & Barber I. (2009) Individual experience and evolutionary history of predation affect expression of heritable variation in fish personality and morphology. Proc. Roy. Soc. Lond. B. 276, 1285-1293.
  • McDonald, P.G., Kazem, A.J.N., Clarke, M.F. & Wright, J. (2008b) Helping as a signal: does removal of potential audiences influence helper provisioning behaviour in the cooperative bell miner? Behav. Ecol. 19, 1047-1055.
  • Sæther, B-E, Engen, S., Grøtan, V., Bregnballe, T., Both, C. Tryjanowski, P., Leivits, A., Wright, J., Møller, A.P., Visser, M. & Winkel, W. (2008) Forms of density regulation in (quasi-) stationary distributions of population sizes in birds. Oikos 117, 1197-1208.
  • McDonald, P.G. & Wright, J. (2008) Provisioning vocalizations in cooperative bell miner: more than a simple stimulus for nestling begging? Auk, 125, 670-678.
  • Smiseth, P.T., Wright, J. & Kölliker, M. (2008) Parent-offspring conflict and co-adaptation: behavioural ecology meets quantitative genetics. Proc. Roy. Soc. Lond. B. 275, 1823-1830.
  • Pacheco, M.L., McDonald, P.G., Wright, J., Kazem, A.J.N. & Clarke, M.F. (2008) Helper contributions to anti-parasite behaviour in the cooperatively breeding bell miner (Manorina melanophrys). Behav. Ecol. 19, 558-566.
  • Wright, J. & Russell, A.F. (2008) How helpers help: disentangling ecological confounds from the benefits of cooperatively breeding. J. Anim. Ecol. 77, 427-429.
  • Budden, A.E. & Wright, J. (2008) The effect of food availability on begging performance and digestive efficiency. Ibis 150, 234-241.
  • McDonald, P.G., te Marvelde, L., Kazem, A.J.N. & Wright, J. (2008a) Helping as a signal and the effect of a potential audience during provisioning visits in a cooperative bird. Anim. Behav. 75, 1319-1330.
  • Wright, J. (2007) Cooperation theory meets cooperative breeding: exposing some ugly truths about social prestige, reciprocity and group augmentation. Behav. Proc. 76, 142-148.
  • Dingemanse, N.J., Wright, J., Kazem, A.J.N., Thomas, D., Hickling, R., & Dawnay, N. (2007) Behavioural syndromes differ predictably between 12 populations of three-spined stickleback. J. Anim. Ecol. 76, 1128-1138.
  • McDonald, P.G., Kazem, A.J.N. & Wright, J. (2007b) A critical analysis of ‘deceptive’ or ‘false feeding’ behaviour in a cooperative bird: disturbance effects, satiated nestlings or deceit? Behav. Ecol. Sociobiol. 61, 1623-1635.
  • Sæther, B-E, Engen, S., Grøtan, V., Fiedler, W., Matthysen, E., Visser, M.E., Wright, J., Adriansen, F., van Balen, H., Balmer, D., Mainwaring, M.C., McCleery, R.H., Pampus, M. & Winkel, W. (2007) The extended Moran effect and large-scale synchronous fluctuations in the size of great tit and blue tit populations. J. Anim. Ecol. 76: 315-325.
  • McDonald, P.G., Heathcote, C., Clarke, M.F., Wright, J. & Kazem, A.J.N. (2007a) Provisioning calls in the cooperatively breeding bell miner Manorina melanophyrs encode sufficient information for individual discrimination J. Avian Biol. 38, 113-121.
  • McDonald, P.G., Kazem, A.J.N., Clarke, M.F. & Wright, J. (2006) Altruistic acts as indicators of quality in a cooperative bird: are helpers-at-the-nest ‘showing off’? J. Ornith. 147, 9-10 (IOC Abstract).
  • Both C., Sanz J.J., Artemyev A.A., Blaauw B., Cowie R.J., Dekhuijzen A.J., Enemar A., Järvinen A., Nyholm N.E.I., Potti J., Ravussin P.-A., Silverin B., Slater F.M., Sokolov L.V., Visser M.E., Winkel W., Wright J. & Zang H. (2006) Pied flycatchers travelling from Africa to breed in Europe: differential effects of winter and migration conditions on breeding date. Ardea 94: 511-525.
  • Edelaar, P. & Wright, J. (2006) Potential prey make excellent ornithologists: adaptive flexible responses towards avian predation threat in Arabian babblers Turdoides squamiceps living in a migration hot-spot. Ibis 148, 664-671.
  • Wright J., Markman S. & Denney S.M. (2006) Facultative adjustment of pre-fledging mass in swifts. Proc. Roy. Soc. Lond. B. 273, 1895-1900.
  • Yom-Tov, Y., Yom-Tov, S., Wright, J., Thorne, C.J.R. & du Feh, R. (2006) Recent changes in body weight and wing length in some British passerine birds. Oikos 112, 91-101.
  • Budden, A.E. & Wright, J. (2005) Evidence for learning during competitive positioning within the nest: do nestlings use ‘ideal free’ foraging tactics? Behav. Ecol. Sociobiol. 58, 227-236.
  • Leader, N., Wright, J. & Yom-Tov, Y. (2005) Acoustic properties of two urban song dialects in the orange tufted sunbirds (Nectarinia osea). Auk 122, 231-245.
  • Both, C., Artemyev, A.V., Blaauw, B., Cowie, R., Dekhuizen, H., Gustafsson, L., Järvinen, A., Kerimov, A., Metcalfe, N., Potti, J., Ravussin, P-A, Sanz, J.J., Silverin, B., Slater, F., Török, J., Winkel, W., Wright, J., Zang, H., Visser, M.E. (2004) Large-scale geographical variation confirms that climate change causes birds to lay earlier. Proc. Roy. Soc. Lond. B. 271, 1657-1662.
  • Walling, C.A., Dawnay, N., Kazem, A.J.N. & Wright, J. (2004) Predator inspection in sticklebacks (Gasterosteus aculeatus): body size, local predation pressure and cooperation. Behav. Ecol. Sociobiol. 56, 164-170.
  • Markman, S., Pinshow, B., Wright, J. & Kotler, B.P. (2004) Food patch use by parent birds: to gather food for themselves or for their chicks. J. Anim. Ecol. 73, 747-755.
  • Wright, J., Stone, R.E. & Brown, N. (2003) Communal roosts as structured information centres in the raven, Corvus corax. J. Anim. Ecol. 72, 1003-1014.
  • Blount, J.D., Houston, D.C., Møller, A.P. & Wright, J. (2003) Do individual branches of immune defence correlate? A comparative study of scavenging and non-scavenging birds. Oikos. 102, 340-350.
  • Martins, T.L.F., Blakey, J.K. & Wright, J. (2002) Low incidence of extra-pair paternity in the colonially-nesting common swift Apus apus. J. Avian Biol. 33, 441-446.
  • Markman, S., Pinshow, B. & Wright, J. (2002) Manipulation of food resources reveals sex-specific trade-offs between parental self-feeding and offspring care. Proc. Roy. Soc. Lond. B. 269, 1931-1938.
  • Kokko, H., Johnstone, R.A. & Wright, J. (2002) The evolution of parental and alloparental effort in cooperatively breeding groups: when should helpers pay-to-stay? Behav. Ecol. 13, 291-300.
  • Wright, J., Hinde, C., Both, C. & Fazey, I. (2002) Begging signals more than just short-term need: cryptic effects of brood size in the pied flycatcher. Behav. Ecol. Sociobiol. 52, 74-83.
  • Karasov, W.H. & Wright, J. (2002) Nestling Digestive Physiology and Begging. In: The Evolution of Begging: Competition, Cooperation and Communication (Ed by Wright, J. & Leonard ML), pp-199-219, Kluwyer Academic Press, Dordrecht, NL.
  • Wright, J. & Leonard, M.L. (2002) The Evolution of Begging: Competition, Cooperation and Communication. (edited proceeding from Begging Workshop, Gregynog 2000), Kluwyer Academic Press, Dordrecht, NL.
  • Leader, N., Wright, J. & Yom-Tov, Y. (2002) Dialect discrimination by male orange tufted sunbirds (Nectarinia osea): reactions to own versus neighbor dialects. Ethology 108, 367-376.
  • Karasov, W.H., McSwan E.L. & Wright, J. (2001) Begging and digestion by nestling pied flycatchers in relation to short- and long-term nutritional need. Am. Zool. 41, 1490-1490 (SIBC abstract).
  • Wright, J., Berg, E., de Kort, S., Khazin, V. & Maklakov, A. (2001c) Cooperative sentinel behaviour in the Arabian babbler. Anim. Behav. 62, 973-979.
  • Wright, J., Berg, E., de Kort, S., Khazin, V. & Maklakov, A. (2001b) Safe selfish sentinels in a cooperative bird. J. Anim. Ecol. 70, 1070-1079.
  • Wright, J., Maklakov, A. & Khazin, V. (2001a) State-dependent sentinels: an experimental study in the Arabian babbler. Proc. R. Soc. Lond. B. 268, 1-6.
  • Denny, A.J., Wright, J. & Grief, B. (2001) Foraging trail efficiency in the wood ant Formica rufa: is time of the essence? Anim. Behav. 62, 139-146.
  • Budden, A.E. & Wright, J. (2001b) Falling on deaf ears: the adaptive significance of begging in the absence of a parent. Behav. Ecol. Sociobiol. 49, 474-481.
  • Budden, A.E. & Wright, J. (2001a) Begging in Nestling Birds. Current Ornithology 16, 83-118.
  • Budden, A.E. & Wright, J. (2000) Nestling diet, chick growth and breeding success in the southern grey shrike (Lanius meridionalis). Proc. 3rd Int. Shrike Symp. Gdansk 1999. The Ring 22, 165-172.
  • Leader, N., Wright, J. & Yom-Tov, Y. (2000) Microgeographic song variation in the orange-tufted sunbird (Necterinia osea). Behaviour 137, 1613-1637.
  • Yom-Tov, Y., Wright, J. & Both, C. (2000) Intraspecific nest parasitism and nest guarding in the pied flycatcher, Ficedula hypoleuca. Ibis 142, 331-332.
  • Whitehead, S., Wright, J. & Cotton, P.A. (1999) Patterns of winter field use by starlings (Sturnus vulgaris) - how important is prey depletion? Bird Study 46, 289-298.
  • Wright, J. (1999b) Adaptive versus non-adaptive helping in cooperative breeders. Behav. Ecol. Sociobiol. 46, 437-438.
  • Johnstone, R.A., Woodroffe, R., Cant, M.A. & Wright, J. (1999) Reproductive skew in multi-member groups. Am. Nat. 153, 315-331.
  • Wright, J., Parker, P., & Lundy K. (1999) Relatedness and chick-feeding effort in the cooperatively-breeding Arabian babbler. Anim. Behav. 58, 779-785.
  • Wright, J. & Dingemanse, N. (1999) Parents and helpers compensate for experimental changes in the provisioning effort of others in the Arabian babbler. Anim. Behav. 58, 345-350.
  • Cotton, P.A., Wright, J. & Kacelnik, A. (1999) Chick begging strategies in relation to brood hierarchies and hatching asynchrony. Am. Nat. 153, 412-420.
  • Wright, J. (1999a) Altruism as a signal: Zahavi’s alternative to kin selection and reciprocity. J. Avian Biol. 30, 108-115.
  • Markman, S., Pinshow, B. & Wright, J. (1999) Orange-tufted sunbirds do not feed nectar to their chicks. Auk 116, 257-259.
  • Wright, J., Both, C., Cotton, P.A. & Bryant, D. (1998) Quality versus quantity: energetic and nutritional trade-offs in parental provisioning strategies. J. Anim. Ecol. 67, 620-634.
  • Wright, J. (1998c) Paternity and Paternal Care. Chapter In: Birkhead, T.R. and Møller, A.P. Sperm Competition and Sexual Selection. Academic Press, London pp. 117-145.
  • Wright, J. (1998b) Helpers-at-the-nest have the same provisioning rule as parents: experimental evidence from playbacks of chick begging. Behav. Ecol. Sociobiol. 42, 423-429.
  • Wright, J. (1998a) Helping-at-the-nest and group size in the Arabian babbler (Turdoides squamiceps). J. Avian Biol. 29, 105-112.
  • Markman, S., Pinshow, B. & Wright, J. (1998) Quality of food eaten by parent sunbirds directly affects their brood-care effort. Ostrich 69, 196 (IOC abstract).
  • Wright, J. (1997) Helping-at-the-nest in Arabian babblers: signalling social status or sensible investment in chicks? Anim. Behav. 54, 1439-1448.
  • Bradbury, R.B., Cotton, P.A., Wright, J., & Griffiths, R. (1997) Nestling sex ratios in the European Starling Sturnus vulgaris. J. Avian Biol. 28, 255-258.
  • Cotton, P.A., Kacelnik, K. & Wright, J. (1996) Chick begging as a signal: are nestlings honest? Behav. Ecol. 7, 178-182.
  • Whitehead, S., Wright, J. & Cotton, P.A. (1996) Measuring the impact of parental foraging by starlings, Sturnus vulgaris, on soil invertebrates: an exclosure experiment. Oikos 76, 511-521.
  • Markman, S., Yom-Tov, Y. & Wright, J. (1996) The effect of male removal on female parental care in the Orange-Tufted Sunbird. Anim. Behav. 52, 437-444.
  • Markman, S., Yom-Tov, Y. & Wright, J. (1995) Male parental care in the Orange-Tufted Sunbird: behavioural adjustment in provisioning and nest guarding effort. Anim. Behav. 50, 655-669.
  • Whitehead, S., Wright, J. & Cotton, P.A. (1995) Winter field use by the European Starling Sturnus vulgaris: habitat preferences and the availability of prey. J. Avian Biol. 26., 193-202.
  • Kacelnik, K., Cotton, P.A., Stirling, L. & Wright, J. (1995) Food allocation among nestling starlings: sibling competition and the scope of parental choice. Proc. Roy. Soc. Lond. B. 259, 259-263.
  • Wright, J. & Cotton, P.A. (1994b) Song versus food: trade-off decisions in a breeding colony of European starlings (Sturnus vulgaris). Bird Study 41, 95-101.
  • Wright, J. & Cotton, P.A. (1994a) Experimentally-induced sex differences in parental care: an effect of certainty of paternity? Anim. Behav. 47, 1311-1322.
  • Yom-Tov, Y. & Wright, J. (1993) The effect of heating nestboxes on egg laying in the Blue Tit, Parus caeruleus. Auk 110, 95-99.
  • Martins, T.L.F. & Wright, J. (1993c) Patterns of food allocation between parent & young under differing weather conditions in the Common Swift (Apus apus). Avocetta 17, 147-156.
  • Martins, T.L.F. & Wright, J. (1993b) The cost of reproduction and the allocation of food between parent and young in the common swift (Apus apus). Behav. Ecol. 4, 213-223.
  • Martins, T.L.F. & Wright, J. (1993a) Brood reduction in response to manipulated brood sizes in the common swift (Apus apus). Behav. Ecol. Sociobiol. 32, 61-70.
  • Wright, J. (1992) Certainty of paternity and parental care. Anim. Behav. 44, 380-381.
  • Wright, J. & Cuthill, I. (1992) Monogamy in the European starling. Behaviour 120, 262-285.
  • Wright, J. & Cuthill, I. (1990b) Biparental care: short term manipulation of partner contribution and brood size in the starling Sturnus vulgaris. Behav. Ecol. 1, 116-124.
  • Wright, J. & Cuthill, I. (1990a) Manipulation of sex differences in parental care: the effect of brood size. Anim. Behav. 40, 462-471.
  • Wright, J. & Cuthill, I. (1989) Manipulation of sex differences in parental care. Behav. Ecol. Sociobiol. 25, 171-181.
  • Hailey, A., Wright, J., & Steer, E. (1988) Population ecology and conservation of tortoises: the effects of disturbance. Herp. J. 1, 294-301.
  • Wright, J., Steer, E., & Hailey, A. (1988) Habitat separation in tortoises and the consequences for activity and thermoregulation. Can. J. Zool. 66, 1537-1544.




Professor in Biology (Ethology), NTNU, Norway (2003-present)

Senior Lecturer in Animal Ecology UWB, UK (2002-2003)

Lecturer in Animal Ecology, UWB, UK (1996-2002)

NERC Research Fellowship, Cambridge, UK (1993-1996)

Post-doc Research Assistant, Oxford, UK (1990-1993)

D.Phil (Oxon) Zoology, Oxford, UK (1986-1990)

BSc (Hons) Ecology, 1st Class, UEA, UK (1983-1986)


Har produsert 8 doktorgradstudenter, 35 masterstudenter og 51 honoursstudenter, mer enn 100 vitenskapelige artikler, >65 seminarer og  >70 internasjonal conferense presentasjoner, bevilget >5mil NOK i forskningsstøtte, redaktør for Ethology, J.Anim. Ecol. og Phil. Trans. Roy. Soc. Lond. B., anmelder for forskningråd i fem nasjoner, samt artikler for alle internasjonale tidsskifter i etologi, evolusjon og økologi.














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