| In a recent revision (Soleglad & Sissom, 2001), the family Scorpiopidae and
the chactid genus Chactopsis were included into the Euscorpiidae. This was done after
a very thorough phylogenetic analysis of the two families.
After the revision, this family now includes 14 genera and 173 species [Updated 13.10.20].
Euscorpiidae are widespread
in central and southern Europe, and also found in Africa (Mediterranean coast), North
America (Mexico), Central America (Guatemala), South America (Brazil, Peru, Venezuela),
Asia (west, central, south and southeast).
One species has become established in some parts of southern
England. The forefathers of these scorpions probably came with
merchandise to the harbors, and have succeeded in surviving in
sheltered places.
Some species of this family have been reported in
captivity, especially members of the genus Euscorpius.
Some members of the genus Scorpiops have been
reported to occur in captivity, but many of these might be
misidentified members of the scorpionid genus
Heterometrus.
The members of the genus Euscorpius are under taxonomic
investigations by professor Victor Fet and other scientists.
Several important changes are expected, and these will be
published in The Scorpion Files as soon as they are
official.
Online
identification key for the Euscorpius genus. This key is now outdated!
Euscorpiids are harmless scorpions which possess no threat
to healthy humans.
Some
habitat pictures for E. carpathicus in Italy.
Some
habitat pictures for E. flavicaudis in England.
Soleglad, M. E. & Sissom, W. D. 2001: Phylogeny of the family Euscorpiidae
Laurie, 1896: a major revision. 25-111. In V. Fet & P. A. Selden (eds.). Scorpions 2001. In memoriam
Gary A. Polis. British Arachnological Society. Burnham Beeches, Bucks. xi + 404 pp.
Genus Chactopsis has been transfered back to Chactidae based on the decisions in Lourenco, 2003 and Lourenco et al., 2011.
Thanks to Dr. Victor Fet and Dr. Benjamin Gantenbein for
keeping me updated on Euscorpius taxonomy!
Euscorpius flavicaudis photo (left)
by Jan Ove Rein (C).
Scorpiops longimanus photo (right) by Dr.
Wilson Lourenço (C) and The Journal of Venomous Animals and Toxins/
CEVAP (C)
SPECIES
FILES:
This list of genera and
species is based on Fet
et al.(2000) and Soleglad & Sissom, 2001. September 2019: Subspecies are included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000) |
Alloscorpiops Vachon, 1980* (7)
A. anthracinus (Simon, 1887)
A. calmonti Lourenço, 2013*
A. citadelle Kovarik, 2013*
A. lindstroemii (Thorell, 1889)
A. troglodytes Lourenço & Pham, 2015*
A. viktoriae Lourenço & Koulic, 2018*
A. wongpromi Kovarik, Soleglad & Kosulic, 2013* |
Alpiscorpius Gantenbein et al., 1999* (15)
A. alpha (Caporiaco, 1950)*
A. beroni (Fet, 2000)*
A. beta (Di Caporiacco, 1950)*
A. delta Kovarik, Stundlova, Fet & Stahlavsky, 2019*
A. gamma (Caporiaco, 1950)*
A. germanus (C.L. Koch, 1837)
A. kappa Kovarik, Stundlova, Fet & Stahlavsky, 2019*
A. lambda Kovarik, Stundlova, Fet & Stahlavsky, 2019*
A. mingrelicus (Kessler, 1874)
A. mingrelicus mingrelicus (Kessler, 1874)
A. mingrelicus caporiaccoi (Bonacina, 1980)
A. mingrelicus dinaricus (Caporiacco, 1950)
A. omega Kovarik, Stundlova, Fet & Stahlavsky, 2019*
A. omikron Kovarik, Stundlova, Fet & Stahlavsky, 2019*
A. phrygius (Bonacina, 1980)*
A. sigma Kovarik, Stundlova, Fet & Stahlavsky, 2019*
A. uludagensis (Lacroix, 1995)*
A. ypsilon Kovarik, Stundlova, Fet & Stahlavsky, 2019*
|
Dasyscorpiops
Vachon, 1974* (1)
D. grandjeani Vachon, 1974
|
Euscorpiops
Vachon, 1980* (28)
E. alexandreanneorum Lourenço, 2013*
E. artemisae Kovarík, Kosulic, Stahlavsky, Dongkhamfu & Wongprom 2015*
E. asthenurus (Pocock, 1900)
E. beccaloniae Kovarík, 2005*
E. bhutanensis (Tikader & Bastawade, 1983)
E. binghamii (Pocock, 1893)
E. cavernicola Lourenço & Pham, 2013*
E. chiangmai Lourenço, 2019*
E. dakrong Lourenço & Pham, 2014*
E. kaftani (Kovarík, 1993)
E. kamengensis Bastawade, 2006*
E. karschi Qi, Zhu & Lourenço, 2005*
E. kubani Kovarík, 2004*
E. lii Di & Qiao, 2020*
E. longimanus (Pocock, 1893)
E. montanus (Karsch, 1879)*
E. neradi Kovarik, Pliskova & Stahlavsky, 2013*
E. novaki Kovarík, 2005*
E. orioni Kovarík, Kosulic, Stahlavsky, Dongkhamfu & Wongprom 2015*
E. problematicus (Kovarik, 2000)*
E. puerensis Di, Wu, Cao, Xiao & Li, 2010*
E. sejnai (Kovarik, 2000)*
E. shidian Qi, Zhu & Lourenço, 2005*
E. thaomischorum Kovarík, 2012*
E. vachoni Qi, Zhu & Lourenço, 2005*
E. validus Di, Cao, Wu & Li, 2010*
E. xui Sun & Zhu, 2010*
E. yangi Zhu, Zhang & Lourenço, 2007*
E. zhangshuyuani Ythier, 2019*
|
Euscorpius Thorell, 1876 (62)
E. aladaglarensis Tropea & Yagmur, 2016*
E. alanyaensis Tropea, Yagmur, Parmakelis & Kunt, 2016*
E. altadonnai Tropea, 2017*
E. amorgensis Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2017*
E. aquilejensis (C. L. Koch, 1837)*
E. arikani Yagmur & Tropea, 2015*
E. avcii Tropea, Yagmur, Koc, Yesilyurt & Rossi 2012*
E. balearicus Caporiacco, 1950*
E. biokovensis Tropea & Ozimec, 2020*
E. birulai Fet, Soleglad, Parmakelis, Kotsakiozi & Stathi, 2014*
E. bonacinai Kovarik & Stahlavsky, 2020*
E. borovaglavaensis Tropea, 2015*
E. calabriae Di Caporiacco, 1950*
E. candiota Birula, 1903*
E. canestrinii (Fanzago, 1872)*
E. carpathicus (Linnaeus, 1767)
E. carpathicus carpathicus (Linnaeus, 1767)
E. carpathicus banaticus (C. L. Koch, 1841)
E. celanus Tropea, 2012*
E. ciliciensis Birula, 1898*
E. concinnus (C. L. Koch, 1837)*
E. corcyraeus Tropea & Rossi, 2012*
E. croaticus Di Caporiacco, 1950*
E. curcici Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2017*
E. deltshevi Fet, Graham, Webber & Blagoev, 2014*
E. drenskii Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2015*
E. erymanthius Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2013*
E. eskisehirensis Tropea & Yagmur, 2015*
E. feti Tropea, 2013*
E. garganicus Caporiacco, 1950*
E. garganicus garganicus Caporiacco, 1950
E. garganicus molisanus Tropea, 2017
E. giachinoi Tropea & Fet, 2015*
E. gocmeni Tropea, Yagmur & Yesilyurt, 2014*
E. hadzii Caporiacco, 1950*
E. hakani Tropea & Yagmur, 2016*
E. honazicus Tropea, Yagmur, Karampatsou, Parmakelis & Yesilyurt, 2016*
E. hyblaeus Tropea, 2016*
E. idaeus Yagmur & Tropea, 2017*
E. italicus (Herbst, 1800)
E. kabateki Kovarik & Stahlavsky, 2020*
E. janstai Kovarik & Stahlavsky, 2020*
E. kinzelbachi Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2014*
E. koci Tropea & Yagmur, 2015*
E. koschewnikowi Birula, 1900*
E. kritscheri Fet, Soleglad, Parmakelis, Kotsakiozi & Stathi, 2013*
E. lycius Yagmur, Tropea & Yesilyurt, 2013*
E. mylonasi Fet, Soleglad, Parmakelis, Kotsakiozi & Stathi, 2014*
E. naupliensis (C. L. Koch, 1837)*
E. oglasae Di Caporiacco, 1950*
E. ossae Di Capriacco, 1950*
E. parthenopeius Tropea, Parmakelis, Sziszkosz, Balanika & Bouderka, 2014*
E. popovi Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2015*
E. sadileki Kovarik & Stahlavsky, 2020*
E. salentinus Tropea, 2017*
E. scaber Birula, 1900*
E. scheraboni Kovarik & Stahlavsky, 2020*
E. sicanus (C. L. Koch, 1837)*
E. solegladi Fet, Graham, Webber & Blagoev, 2014*
E. stahlavskyi Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2014*
E. studentium Karaman,2020*
E. sultanensis Tropea & Yagmur, 2016*
E. tauricus (C. L. Koch, 1837)*
E. tergestinus (C.L. Koch, 1837)
E. vailatii Tropea & Fet, 2015*
E. vignai Tropea, Fet, Parmakelis, Kotsakiozi & Stathi, 2014*
E. yagmuri Kovarik, Fet & Soleglad, 2014*
|
Megacormus Karsch, 1881 (6)
M. franckei Kovarik, 2019*
M. gertschi Díaz Najera, 1966
M. granosus (Gervais, 1843)
M. grubbsi Sissom, 1994
M. segmentatus Pocock, 1900
M. xichu Gonzalez-Santillan, Gonzalez-Ruiz & Escobedo-Morales, 2017*
|
Neoscorpiops Vachon, 1980* (5)
N. deccanensis (Tikader & Bastawade, 1978)
N. maharashtraensis Mirza, Sanap & Upadhye, 2014
N. phaltanensis Sulakhe, Sayyed, Deshpande, Dandekar, Padhye & Bastawade, 2020*
N. satarensis (Pocock, 1900)
N. tenuicauda (Pocock, 1894) |
|
Parascorpiops
Banks, 1928* (1)
P. montanus Banks, 1928
|
Plesiochactas
Pocock, 1900 (3)
P. dilutus (Karsch, 1881)
P. mitchelli Soleglad, 1976
P. vasquezi Trujillo & Armas, 2012* |
Plethoscorpiops Lourenço, 2017* (1)
P. profusus Lourenço, 2017*
| Scorpiops Peters,
1861* (39)
S. afghanus Lourenço & Qi, 2006*
S. atomatus Qi, Zhu & Lourenço, 2005*
S. braunwalderi Kovarik, 2000*
S. dastychi Kovarik, 2000*
S. demisi Kovarík, 2005*
S. farkaci Kovarík, 1993
S. feti Kovarik, 2000*
S. furai Kovarík, 2020*
S. grosseri Kovarík, 2020*
S. hardwickii (Gervais, 1843)
S. harmsi Kovarík, 2020*
S. hofereki Kovarík, 2020*
S. ingens Yin, Zhang, Pan, Li & Di, 2015 *
S. irenae Kovarík, 1994
S. jendeki Kovarik, 1994*
S. kejvali Kovarík, 2020*
S. kraepelini Lourenço, 1998
S. langxian Qi, Zhu & Lourenço, 2005*
S. leptochirus Pocock, 1893
S. lhasa Di & Zhu, 2009*
S. lindbergi (Vachon, 1980)*
S. luridus Qi, Zhu & Lourenço, 2005*
S. margerisonae Kovarik, 2000*
S. oligotrichus Fage, 1933
S. pachmarhicus Bastawade, 1992
S. pakistanus Kovarik & Ahmed, 2009*
S. petersii Pocock, 1893
S. pococki Qi, Zhu & Lourenço, 2005*
S. pseudomontanus Kovarik & Ahmed, 2009*
S. rohtangensis Mani, 1959
S. songi Di & Qiao, 2020*
S. spitiensis Zambre, Sanap & Mirza, 2014*
S. taxkorgan Lourenço, 2018
S. tibetanus Hirst, 1911
S. tryznai Kovarík, 2020*
S. vonwicki Birula, 1913*
S. wrzecionkoi Kovarík, 2020*
S. yagmuri Kovarík, 2020*
S. zubairahmedi Kovarik, 2009*
S. zubairi Kovarík, 2020*
|
Tetratrichobothrius Birula, 1917 (1)
T. flavicaudis (DeGeer, 1778)
|
|
Troglocormus
Francke, 1981 (2)
T. ciego
Francke, 1981
T. willis Francke, 1981
|
Vietscorpiops (Lourenço and Pham, 2015) (1)
V. dentidactylus (Lourenço and Pham, 2015)
|
| |
