BUTHIDAE
C.L. Koch, 1837

With 96 genera (one extinct and not listed below) and 1232 species [Updated 06.04.21], this family is the largest of the scorpion families. This family is widespread around the world (not found in Antarctica and New Zealand), and are found in tropical, subtropical and partly in temperate habitats.

Volschenk, Mattoni & Prendini (2008) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus and Neoprotobuthus) and 15 species are all transfered to Buthidae. August 2019: The family Microcharmidae has been resurrected by Lourenco et al., 2019 and belinging taxa have been transferred back from Buthidae Information about the family Microcharmidae.

Kovarik (2007) has published a revision of the genus Hottentotta. Note that the medical important Mesobuthus tamulus and two other Asian Mesobuthus have been transfered to Hottentotta. Kovarik's revision can be downloaded HERE (browse down to issue 58).

Paraorthochirus has been synonymized with Orthochirus by Navidpour et al., 2008).

The members of this family are small and medium sized scorpions with a triangular sternum (some genera have a more pentagonal sternum). In many genera (e.g., Androctonus, Apistobuthus, Parabuthus, and other Old World genera) the cauda is usually strong and powerful and the pedipalps are often very slender whereas other genera have elongate appendages, especially the males (e.g., Centruroides, Lychas, Isometrus and related genera). Many species are yellow or brown (or variations of these colors), but black forms are also present. Some species have quite intricate and attractive patterns (e.g., Lychas and Isometrus) and colours (e.g., Centruroides and Uroplectes). Sizes range from about 20 mm (like Microtityus and Microbuthus) to over 120 mm (as in Androctonus, some Centruroides, Apistobuthus and others).

An indentification key to the genus Compsobuthus was published by Kovarik (2012) [Free full text].

Several species in this family are highly toxic, but fewer than 20 can be lethal to man. The most notorious species are found in the genus Androctonus, Centruroides, Hottentotta, Leiurus, Parabuthus and Tityus. See Scorpions of medical importance page for further information.

Unlike most other scorpion families, the Buthidae are of both scientific and public interest due to their medical importance in many parts of the world. Several species have also made their way into the hobby. Many of these are safe to be kept by experienced collectors, but the most dangerous species should be avoided.

Androctonus crassicauda habitat picture from Kuwait are located HERE.

Photo of Parabuthus leiosoma (left) by Jan Ove Rein (C)
Photo of Hottentotta sp. (right) by Jan Ove Rein (C)

SPECIES FILES:
This list of genera and species is based on Fet et al.(2000). September 2019: Subspecies are included in the list. I try to update the list as additions and changes are published. * denotes changes after Fet et al. (2000). I will be grateful for information about new development in the taxonomy of this family. For information about synonyms and bibliographies, see Fet et al. (2000)

-
Aegaeobuthus Kovarik, 2019* (4)
A. cyprius (Gantenbein & Kropf, 2000)*
A. gallianoi (Ythier, 2018)*
A. gibbosus (Brulle, 1832)
A. nigrocinctus (Ehrenberg, 1828)*
A. nigrocinctus bishri (Lourenco, 2020)*
Afrolychas Kovarik, 2019* (2)
A. braueri (Kraepelin, 1896)
A. burdoi (Simon, 1882)
Afroisometrus Kovarik, 1997 (1)
A. minshullae (Fitzpatrick, 1994)
Akentrobuthus Lamoral, 1976 (2)
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976
Alayotityus Armas, 1973 (8)
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A. juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas, 1973

Ananteris Thorell, 1891 (92)
A. arcadioi Botero-Trujillo, 2008*
A. ashaninka Kovarik, Teruel, Lowe & Friedrich, 2015*
A. ashmolei Lourenço, 1981
A. asuncionensis González-Sponga, 2006*
A. balzanii Thorell, 1891
A. barinensis González-Sponga, 2006*
A. bernabei Giupponi, Vasconcelos & Lourenço, 2009*
A. bianchinii Lourenço, Aguiar-Neto & Limeira-de-Oliveira, 2009*
A. bonito Lourenço, 2012*
A. cachimboensis Lourenço, Motta & da Silva, 2006*
A. camacan Lourenço, Giupponi & Leguin 2013*
A. canalera Miranda & Armas, 2020*
A. capayaensis González-Sponga, 2006*
A. caracensis González-Sponga, 2006*
A. carrasco Lourenço & Motta, 2019*
A. catuaroi González-Sponga, 2006*
A. caucaguitensis González-Sponga, 2006*
A. chagasi Giupponi, Vasconcelos & Lourenço, 2009*
A. charlescorfieldi Lourenço, 2001*
A. chirimakei González-Sponga, 2006*
A. cisandinus Lourenço, 2015*
A. claviformis González-Sponga, 2006*
A. coineaui Lourenço, 1982
A. columbiana Lourenço, 1991
A. cryptozoicus Lourenço, 2005*
A. cumbensis González-Sponga, 2006*
A. curariensis González-Sponga, 2006*
A. cussinii Borelli, 1910
A. dacostai Ythier, Chevalier & Lourenço, 2020*
A. dekeyseri Lourenço, 1982
A. deniseae Lourenço, 1997
A. desiderio Lourenço, Giupponi & Leguin 2013*
A. diegorojasi Rojas-Runjaic, 2005*
A. dorae Botero-Trujillo, 2008*
A. ehrlichi Lourenço, 1994
A. elguapoi González-Sponga, 2006*
A. elisabethae Lourenço, 2003*
A. evellynae Lourenço, 2004*
A. faguasi Botero-Trujillo, 2009*
A. festae Borelli, 1899
A. franckei Lourenço, 1982
A. gorgonae Lourenço & Flórez, 1989
A. guiripaensis González-Sponga, 2006*
A. guyanensis Lourenço & Monod, 1999*
A. infuscata Lourenço, Giupponi & Leguin 2013*
A. inoae González-Sponga, 2006*
A. intermedia Lourenço, 2012*
A. kalina Ythier, 2018*
A. kuryi Giupponi, Vasconcelos & Lourenço, 2009*
A. leilae Lourenço, 1999*
A. luciae Lourenço, 1984
A. madeirensis Lourenço & Duhem, 2010*
A. mamilihpan Ythier, Chevalier & Lourenço, 2020*
A. maniapurensis González-Sponga, 2006*
A. maranhensis Lourenço, 1987
A. mariaelenae Lourenço, 1999*
A. mariaterezae Lourenço, 1982
A. mauryi Lourenço, 1982
A. meridana González-Sponga, 2006*
A. michaelae Lourenço, 2013*
A. myriamae Botero-Trujillo, 2007*
A. nairae Lourenço, 2004*
A. norae González-Sponga, 2006*
A. ochoai Botero-Trujillo & Florez D, 2011*
A. otavianoi Lira, Pordeus & Ribeiro de Albuquerque, 2017
A. palmari Botero-Trujillo & Noriega, 2011*
A. paoensis González-Sponga, 2006*
A. paracotoensis González-Sponga, 2006*
A. pierrekondre Lourenço, Chevalier, Gangadin & Ythier, 2020*
A. plataensis González-Sponga, 2006*
A. platnicki Lourenço, 1993
A. polleti Lourenço, 2016*
A. principalis González-Sponga, 2006*
A. pydanieli Lourenço, 1982
A. riocaurensis González-Sponga, 2006*
A. riochicoi González-Sponga, 2006*
A. riomachensis Rojas-Runjaic, Portillo-Quintero & Borges, 2008*
A. roraima Lourenço & Duhem, 2010*
A. sabineae Lourenço, 2001*
A. sanchezi González-Sponga, 2006*
A. sepulvedai González-Sponga, 2006*
A. singularis González-Sponga, 2006*
A. sipilili Ythier, Chevalier & Lourenço, 2020*
A. solimariae Botero-Trujillo & Florez D, 2011*
A. surinamensis Lourenço, 2012*
A. tresor Ythier, Chevalier & Lourenço, 2020*
A. terueli Kovarik, 2006*
A. tolimana Teruel & Garcia, 2007*
A. turumbanensis Gonzales-Sponga, 1980
A. venezuelensis Gonzales-Sponga, 1972
A. volschenki Botero-Trujillo, 2009*
A. zuliana González-Sponga, 2006*

Ananteroides Borelli, 1911* (2)
A. feae Borelli, 1911*
A. inexpectatus Lourenço, 2013*
 
Androctonus Ehrenberg, 1828 (29)
A. aeneas C. L. Koch, 1839*
A. afghanus Lourenço & Qi, 2006*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi (Audouin,1826)
A. amoreuxi amoreuxi (Audouin,1826)
A. amoreuxi levyi Fet, 1997
A. australis (Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. barbouri (Werner, 1932)*
A. bicolor Ehrenberg, 1828
A. bicolor bicolor Ehrenberg, 1828
A. bicolor longecarinatus (Caporiacco, 1932) [nomen dubium]
A. cholistanus Kovarik & Ahmed, 2013*
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. donairei Rossi, 2015*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Ythier & Leguin, 2009*
A. pallidus Lourenço, Duhem & Cloudsley-Thompson, 2012*
A. robustus Kovarik & Ahmed, 2013*
A. santi Lourenço, 2015*
A. sergenti Vachon, 1948
A. simonettai Rossi, 2015*
A. tenuissimus Teruel, Kovarik & Turiel, 2013*
A. tigrai Lourenço, Rossi & Sadine 2015*
A. togolensis Lourenço, 2008*
A. tropeai Rossi, 2015*
Anomalobuthus Kraepelin, 1900 (6)
A. krivochatskyi Teruel, Kovarik & Fet, 2018*
A. lowei Teruel, Kovarik & Fet, 2018*
A. pavlovskyi Teruel, Kovarik & Fet, 2018*
A. rickmersi Kraepelin, 1900
A. talebii Teruel, Kovarik, Navidpour & Fet, 2014*
A. zarudnyi (Birula, 1911)*
Apistobuthus Finnegan, 1932 (2)
A. pterygocercus Finnegan, 1932
A. susanae Lourenço, 1998
Australobuthus Locket, 1990 (1)
A. xerolimniorum Locket, 1990
Babycurus Karsch, 1886 (14)
B. ansorgei Hirst, 1911
B. brignolii Lourenço & Rossi, 2017* [nomen dubium]
B. buettneri Karsch, 1886
B. buettneri buettneri Karsch, 1886
B. buettneri savanicola Lourenco, Bruehmueller Ramos & Cloudsley-Thompson, 2005
B. centrurimorphus Karsch, 1886*
B. dunlopi Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. pictus Pocock, 1896*
B. solegladi Lourenço, 2005*
B. taramassoi Borelli, 1919*
B. wituensis Kraepelin, 1913
Baloorthochirus Kovarik, 1996 (1)
B. becvari Kovarik, 1996
Barbaracurus Kovarik, Lowe & Stahlavsky, 2018 (10)
B. feti Kovarik, Lowe &Stahlavsky & Hurre, 2019*
B. exquisitus (Lowe, 2000)*
B. prudenti (Lourenço, 2013)*
B. somalicus (Hirst, 1907)*
B. sofomarensis (Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015)*
B. subpunctatus (Borelli, 1925)*
B. ugartei (Kovarik, 2000)*
B. winklerorum Kovarik, Lowe &Stahlavsky, 2018*
B. yemenensis Kovarik, Lowe &Stahlavsky, 2018*
B. zambonellii (Borelli, 1902)*
Birulatus Vachon, 1974 (4)
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
B. jordanensis Lourenço, Al-Saraireh, Afifeh, Baker, Bader-Katbeh & Amr, 2021*
Buthacus Birula, 1908 (23)
B. agarwali Zambre & Lourenço, 2010*
B. ahaggar Lourenço, Kourim & Sadine, 2017*
B. arenicola (Simon, 1885)
B. arenicola arenicola (Simon, 1885)
B. arenicola maroccanus Lourenco, 2006
B. birulai Lourenço, 2006*
B. clevai Lourenço, 2001*
B. elmenia Lourenço & Sadine, 2017*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (nomen dubium)
B. golovatchi Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. leptochelys (Ehrenberg, 1829)
B. leptochelys leptochelys (Ehrenberg, 1829)
B. leptochelys granosus Borelli, 1929
B. leptochelys nitzani Levy, Amitai & Shulov, 1973
B. macrocentrus (Ehrenberg, 1828)*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. occidentalis Vachon, 1953*
B. pakistanensis Lourenço & Qi, 2006*
B. samiae Lourenço & Sadine, 2015*
B. spinatus Lourenço, Bissati & Sadine, 2016*
B. stockmanni Kovarik, Lowe & Stahlavsky, 2016*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. williamsi Lourenço & Leguin, 2009*
B. ziegleri Lourenço, 2000*
Butheoloides Hirst, 1925 (19)
B. annieae Lourenço, 1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. granulatus Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. grosseri Kovarik, 2016*
B. hirsti Lourenço, 1996
B. littoralis Lourenço, Touloun & Boumezzough, 2011*
B. maroccanus Hirst, 1925
B. milloti Vachon, 1948
B. monodi Vachon, 1950
B. nuer Kovarik, 2015*
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. savanicola Lourenço, 2013*
B. schwendingeri Lourenço, 2002*
B. slimanii Lourenço, 2010*
B. vanderberghi Lourenço, 2015*
B. wilsoni Lourenço, 1995
Butheolus Simon, 1882 (6)
B. andersoni (Pocock, 1895)*
B. gallagheri Vachon, 1980
B. hallani Lourenço & Rossi, 2017*
B. harrisoni Lowe, 2018*
B. thalassinus Simon, 1882
B. villosus Hendrixson, 2006*
Buthiscus Birula, 1905 (1)
B. bicalcaratus Birula, 1905
Buthoscorpio Werner, 1936 (5)
B. chinnarensis Aswathi, Sureshan & Lourenço, 2015*
B. indicus Lourenço, 2012*
B. politus (Pocock, 1899)
B. rayalensis Javed, Rao, Mirza, Sanap & Tampal, 2010*
B. sarasinorum (Karsch, 1891)
NB! B. jinnahii Amir, Kamaluddin & Jabbar, 2005* and B. rahmatii Amir, Kamaluddin & Jabbar, 2005* was described in the non-valid genus Stenochirus by the authors. I have placed them in Buthoscorpio, which is the valid name for Stenochirus, but formally this has to be done in a scientific journal before it is valid. Javed et al., 2010 have studied the descriptions of these two species and concluded that they do not belong to the genus Buthoscorpio. This decsision was also supported by Lourenco, 2012. The two taxa are now Buthidae incertae sedis and are not included in the total number species in Buthidae.
Buthus Leach, 1815 (66)
B. adrianae Rossi, 2013
B. alacanti Teruel & Turiel 2020*
B. albengai Lourenço, 2003*
B. amri Lourenço, Yagmur & Duhem 2010*
B. apiatus Lourenço, El Bouhissi & Sadine, 2020*
B. atlantis Pocock, 1889
B. aures Lourenço & Sadine 2016*
B. awashensis Kovarik, 2011*
B. baeticus Teruel & Turiel 2020*
B. barcaeus Birula, 1909*
B. berberensis Pocock, 1900*
B. bonito Lourenço & Geniez, 2005*
B. boumalenii Touloun & Boumezzough, 2011*
B. boussaadi Lourenço, Chichi & Sadine, 2018*
B. brignolii Lourenço, 2003*
B. centroafricanus Lourenço, 2016*
B. chambiensis Kovarik, 2006*
B. confluens Lourenço, Touloun & Boumezzough, 2012*
B. danyii Rossi, 2017*
B. delafuentei Teruel & Turiel 2020*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. duprei Rossi & Tropea, 2016*
B. egyptiensis Lourenço, 2012*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. elongatus Rossi, 2012*
B. garcialorcai Teruel & Turiel 2020*
B. hassanini Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. ibericus Lourenço & Vachon, 2004*
B. intermedius (Ehrenberg, 1829)*
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. israelis Shulov & Amitai, 1959*
B. jianxinae Lourenço, 2005*
B. karoraensis Rossi & Tropea, 2016*
B. kunti Yagmur, Koc & Lourenço, 2011*
B. labuschagnei Lourenço, 2015*
B. lienhardi Lourenço, 2003*
B. lourencoi Rossi, Tropea & Yagmur, 2013*
B. malhommei Vachon, 1949*
B. manchego Teruel & Turiel 2020*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. nabataeus Lourenço, Abu Afifeh & Al-Saraireh, 2021*
B. nigrovesiculosus Hirst, 1925*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. orientalis Lourenço & Simon, 2012*
B. oudjanii Lourenço, 2017*
B. paris (C. L. Koch, 1839)*
B. parroti Vachon, 1949*
B. pococki Kovarik, Stahlavsky & Elmi, 2020*
B. prudenti Lourenço & Leguin, 2012*
B. pusillus Lourenço, 2013*
B. rochati Lourenço, 2003*
B. saharicus Sadine, Bissati & Lourenço, 2015*
B. serrano Teruel & Turiel 2020*
B. somalilandus Kovarik, Stahlavsky & Elmi, 2020*
B. tassili Lourenço, 2002*
B. trinacrius Lourenço & Rossi, 2013*+
+Probably extinct.
B. tunetanus (Herbst, 1800)*
B. yemenensis Lourenço, 2008*
B. zeylensis Pocock, 1900*
Centruroides Marx, 1890 (93)
C. alayoni Armas, 1999*
C. altagraciae Teruel, de Armas & Kovarik, 2015*
C. anchorellus Armas, 1976
C. arctimanus Armas, 1976
C. baergi Hoffmann, 1932*
C. balsasensis Ponce & Francke, 2004*
C. bani Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis (Pocock, 1898)
C. bertholdii (Thorell, 1876)
C. bicolor (Pocock, 1898)
C. bonito Quijano-Ravell, Teruel & Ponce-Saavedra, 2016*
C. caral Armas & Trujillo, 2013
C. caribbeanus Teruel & Myers, 2017*
C. chamela Ponce-Saavedra & Francke, 2011*
C. chamulaensis Hoffmann, 1932
C. chiapanensis Hoffmann, 1932
C. edwardsii (Gervais, 1843)*
C. elegans (Thorell, 1876)
C. elegans elegans (Thorell, 1876)
C. elegans edentulus Werner, 1939
C. elegans insularis Pocock, 1902
C. exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul (Meise, 1933)
C. fallassisimus Armas & Trujillo, 2010*
C. farri Armas, 1976
C. flavopictus (Pocock, 1898)
C. flavopictus flavopictus (Pocock, 1898)
C. flavopictus meridionalis Hoffmann, 1932
C. franckei Santibanez-Lopez & Contreras-Felix, 2013*
C. fulvipes (Pocock, 1898)
C. galano Teruel, 2001*
C. gracilis (Latreille, 1804)
C. granosus (Thorell, 1876)*
C. griseus (C. L. Koch, 1844)
C. guanensis Franganillo, 1930
C. hentzi (Banks, 1900)
C. hirsuticauda Teruel, 2011*
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. huichol Teruel, Ponce-Saavedra & Quijano-Ravell, 2015*
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. ixil Trujillo & Armas, 2016*
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. lauriadnae Ponce-Saavedra & Francke, 2019*
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. lucidus Teruel, Armas & Kovarik, 2015*
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mascota Ponce-Saavedra & Francke, 2011*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus (Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. panamensis Arias & Esposito, 2014*
C. platnicki Armas, 1981
C. pococki Sissom & Francke, 1983
C. polito Teruel, 2007*
C. poncei Teruel, Kovarik, Baldazo-Monsivais & Hoferek, 2015*
C. possanii Gonzalez-Santillan, Galan-Sanchez & Valdez-Velazquez, 2019*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. rodolfoi Santibanez-Lopez & Contreras-Felix, 2013*
C. romeroi Quijano-Ravell, De Armas, Francke & Ponce-Saavedra, 2019*
C. ruana Quijano-Ravell & Ponce-Saavedra, 2016*
C. sanandres Armas, Sarmiento & Florez 2012*
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. simplex (Thorell, 1876)*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. suffusus (Pocock, 1902)
C. suffusus suffusus (Pocock, 1902)
C. tapachulaensis Hoffmann, 1932*
C. tecomanus Hoffmann, 1932*
C. tenuis Thorell, 1876*
C. testaceus (DeGeer, 1778)
C. thorellii (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. villegasi Baldazo-Monsivaiz, Ponce-Saavedra & Flores-Moreno, 2013*
C. vittatus (Say, 1821)
Charmus Karsch, 1879 (5)
C. brignolii Lourenço, 2000*
C. indicus Hirst, 1915
C. laneus Karsch, 1879
C. saradieli Kovarik, Lowe, Ranawana, Hoferek & Jayarathne 2016*
C. sinhagadensis Tikader & Bastawade, 1983
Chaneke Francke, Teruel & Santibanez-Lopez, 2014* (4)
C. aliciae (Armas & Frias, 1998)*
C. baldazoi Kovarik, Teruel & Lowe, 2016*
C. fogoso Francke, Teruel & Santibanez-Lopez, 2014*
C. hofereki Kovarik, Teruel & Lowe, 2016*
Cicileiurus Teruel, 2007* (1)
C. monticola Teruel, 2007*
Cicileus Vachon, 1948 (5)
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
C. hoggarensis Lourenço & Rossi, 2015*
C. latellai Lourenço & Rossi, 2015*
C. montanus Lourenço & Rossi, 2015*
Compsobuthus Vachon, 1949 (49)
C. abyssinicus (Birula, 1903)
C. acutecarinatus (Simon, 1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov, 1973
C. armenicus Lourenço, Leguin & Duhem, 2015*
C. atrostriatus (Pocock, 1897)*
C. berlandi Vachon, 1950
C. birulai Lourenço, Leguin & Duhem, 2010*
C. boucheti Lourenço, Duhem & Cloudsley-Thompson, 2012*
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. eritreaensis Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. garyi Lourenço & Vachon, 2001*
C. humaae Amir, Kamaluddin & Kahn, 2005* (nomen dubium)
C. jakesi Kovarik, 2003*
C. jordanensis Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. klaptoczi (Birula, 1909)
C. krali Kovarik, 2012*
C. levyi Kovarik, 2012*
C. longipalpis Levy, Amitai & Shulov, 1973
C. maidensis Kovarik, 2018*
C. maindroni (Kraepelin, 1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. plutenkoi Kovarik, 2003*
C. polisi Lowe, 2001*
C. rugosulus (Pocock, 1900)
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. sindicus Kovarik & Ahmed, 2011*
C. somalilandus Kovarik, 2012*
C. tassili Lourenço, 2010*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. turieli Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. ullrichi Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. vachoni Sisson, 1994
C. werneri (Birula, 1908)
Congobuthus Lourenço, 1999* (1)
C. fagei Lourenço, 1999*
Darchenia Vachon, 1977 (1)
D. bernadettae Vachon, 1977
Egyptobuthus Lourenço, 1999* (1)
E. vaissadei Lourenço, 1999*
Femtobuthus Lowe, 2010* (1)
F. shutuae Lowe, 2010*
Fetilinia Lowe & Kovarik, 2021* (1)
F. dentator Lowe & Kovarik, 2021*
Gint Kovarik, Lowe, Pliskova & Stahlavsky, 2013* (12)
G. amoudensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. banfasae Kovarik & Lowe, 2019*
G. calviceps (Pocock, 1900)
G. childsi Kovarik, 2018*
G. dabakalo Kovarik & Mazuch, 2015*
G. gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013*
G. gubanensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. insolitus (Borelli, 1925)* (nomen dubium)
G. maidensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. marialuisae Rossi, 2015* (nomen dubium)
G. monicae Rossi, 2015* (nomen dubium)
G. puntlandus Kovarik & Mazuch, 2015*
Grosphus Simon, 1880 (34)
G. ambre Lourenco, Wilmé & Waeber, 2018*
G. ankarafantsika Lourenço, 2003*
G. ankarana Lourenço & Goodman, 2003*
G. annulatus Fage, 1929*
G. bemaraha Lourenco, Wilmé & Waeber, 2018*
G. bicolor Lourenço, 2012*
G. bistriatus Kraepelin, 1900*
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. eliseanneae Lourenço & Wilme, 2016*
G. feti Lourenço, 1996*
G. flavopiceus Kraepelin, 1900*
G. ganzhorni Lourenço, Wilme & Waeber, 2016*
G. grandidieri Kraepelin, 1900*
G. goudoti Lourenço & Goodman, 2006*
G. halleuxi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. hirtus Kraepelin, 1900
G. intertidalis Lourenço, 1999*
G. limbatus (Pocock, 1889)*
G. madagascariensis (Gervais, 1843)
G. magalieae Lourenço, 2014*
G. makay Lourenço & Wilme, 2015*
G. mandena Lourenço, 2005*
G. mahafaliensis Lourenço, Goodman & Ramilijaona, 2004*
G. mavo Lourenço & Rossi 2020*
G. mayottensis Lourenço & Goodman, 2009*
G. olgae Lourenço, 2004*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. rakotoariveloi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. rossii Lourenço, 2013*
G. sabineae Lourenço & Wilme, 2016*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
G. tavaratra Lourenco, Soarimalala & Goodman, 2009*
G. voahangyae Lourenço & Wilme 2015*
G. waeberi Lourenço & Wilme, 2016*
Hemibuthus Pocock, 1900 (2)
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
Hemilychas Hirst, 1911 (1)
H. alexandrinus (Hirst, 1911)
Heteroctenus Pocock, 1893* (9)
H. abudi (Armas & Marcano Fondeur, 1987)*
H. aridicola (Teruel et Armas, 2012)*
H. bonettii (Armas 1999)*
H. garridoi (Armas, 1974)*
H. gibarae (Teruel, 2006)*
H. granulimanus (Teruel, 2006)*
H. junceus (Herbst, 1800)*
H. melloleitaoi (Teruel et Armas, 2006)*
H. princeps (Karsch, 1879)*
Himalayotityobuthus Lourenço, 1997 (2)
H. alejandrae Lourenço, 2003*
H. martensi Lourenço, 1997
Hottentotta Birula, 1908 (58)
H. alticola (Pocock, 1895)
H. alticola alticola (Pocock, 1895)
H. alticola nigrifrons (Pocock, 1900)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004* (nomen dubium)
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. flavidulus Teruel & Rein, 2010*
H. franzwerneri (Birula, 1914)
H. fuscitruncus (Caporiacco, 1936)
H. gambelaensis Kovarik, 2015*
H. gentili (Pallary, 1924)*
H. gibaensis Kovarik, 2015*
H. haudensis Kovarik & Lowe, 2021*
H. hoggarensis Lourenço & Leguin, 2014*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. juliae Kovarik, Yagmur & Fet, 2019*
H. keralaensis Aswhati, Sureshan & Lourenço, 2016*
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. krivokhatskyi Kovarik, Yagmur & Fet, 2019*
H. lacroixi Ythier & Dupre, 2021*
H. lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarik & Kayedi, 2010*
H. mazuchi Kovarik, 2013*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1875)
H. minax minax (L. Koch, 1975)
H. minax occidentalis (Vachon & Stockmann, 1968)
H. minusalta Vachon, 1959*
H. navidpouri Kovarik, Yagmur & Moradi, 2018*
H. nigrimontanus Kovarik & Lowe, 2021*
H. niloticus (Birula, 1928)*
H. novaki Kovarik, 2015*
H. pachyurus (Pocock, 1897)*
H. pellucidus Lowe, 2010*
H. penjabensis (Birula, 1897)*
H. polystictus (Pocock, 1896)
H. reddyi Lourenço, 2015*
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1880)
H. saxinatans Lowe, 2010*
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. sistanensis Kovarik, Yagmur & Moradi, 2018*
H. socotrensis (Pocock, 1889)
H. somalicus Kovarik, 2018*
H. songi (Lourenço, Qi & Zhu, 2005)*
H. sousai Turiel, 2014*
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trailini Kovarik, 2013*
H. trilineatus (Peters, 1861)
H. ugandaensis Kovarik, 2013*
H. vinchu Mirza, Ambekar & Kulkarni, 2019*
H. zagrosensis Kovarik, 1997
Iranobuthus Kovarik, 1997 (1)
I. krali Kovarik, 1997
Ischnotelson Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (2)
I. guanambiensis (Lenarducci, Pinto-da-Rocha & Lucas, 2005)*
I. peruassu Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
Isometroides Keyserling, 1885 (1)
I. vescus (Karsch, 1880)
Isometrus Ehrenberg, 1828 (10)
I. amboli Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. atherii Amir & Kamaluddin, 2008*
I. formosus Pocock, 1894
I. isadensis Tikader & Bastawade, 1983*
I. kovariki Sulakhe, Dandekar, Mukherjee, Pandey, Ketkar, Padhye & Bastawade, 2020*
I. liaqatii Amir & Kamaluddin, 2008*
I. maculatus (DeGeer, 1778)
I. tamhini Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. thurstoni Pocock, 1893
I. thwaitesi Pocock, 1897
Jaguajir Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (3)
J. agamemnon (C. L. Koch, 1839)*
J. pintoi (Mello-Leitão, 1932)*
J. pintoi pintoi (Mello-Leitão, 1932)*
J. pintoi kourouensis (Lourenco, 2008)*
J. rochae (Borelli, 1910)*
Janalychas Kovarik, 2019* (9)
J. albimanus (Henderson, 1919)
J. farkasi (Kovarik, 1997)
J. granulatus Mirza, 2020*
J. heurtaultae (Kovarik, 1997)
J. keralaensis Mirza, 2020*
J. laevifrons (Pocock, 1897)
J. shoplandi (Oates, 1888)
J. srilankensis (Lorenco, 1997)
J. tricarinatus (Simon, 1884)
Karasbergia Hewitt, 1913 (1)
K. methueni Hewitt, 1913
Kraepelinia Vachon, 1974 (1)
K. palpator (Birula, 1903)
Lanzatus Kovarik, 2001* (2)
L. somalicus Kovarik, 2001*
L. somalilandus Kovarik, Lowe & Stahlavsky, 2016*
Leiurus Ehrenberg, 1828 (17)
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. arabicus Lowe, Yagmur & Kovarik, 2014*
L. ater Lourenço, 2019*
L. brachycentrus (Ehrenberg, 1829)*
L. dekeyseri Lourenço, 2020*
L. gubanensis Kovarik & Lowe, 2020*
L. haenggii Lowe, Yagmur & Kovarik, 2014*
L. heberti Lowe, Yagmur & Kovarik, 2014*
L. hebraeus (Birula, 1908)*
L. hoggarensis Lourenço, Kourim & Sadine, 2018*
L. kuwaiti Lourenço, 2020*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. macroctenus Lowe, Yagmur & Kovarik, 2014*
L. quinquestriatus (Ehrenberg, 1828)
L. saharicus Lourenço, 2020*
L. savanicola Lourenço, Qi & Cloudsley-Thompson, 2006*
L. somalicus Lourenço, & Rossi, 2016*
Liobuthus Birula, 1898 (1)
L. kessleri Birula, 1898
Lissothus Vachon, 1948 (3)
L. bernardi Vachon, 1948
L. chaambi Lourenço & Sadine, 2014*
L. occidentalis Vachon, 1950
Lychas C.L. Koch, 1845 (31)
L. aberlenci Lourenço, 2013*
L. armasi Kovarik, 2013*
L. armillatus (Gervais, 1841)*
L. asper (Pocock, 1891)
L. biharensis Tikader & Bastawade, 1983
L. brehieri Lourenço, 2017*
L. buchari Kovarik, 1997
L. cernickai Kovarik, 2013*
L. flavimanus (Thorell, 1888)
L. gravelyi Henderson, 1913
L. hendersoni (Pocock, 1897)
L. hillyardi Kovarik, 1997
L. inexpectatus Lourenço, 2011*
L. kamshetensis Tikader & Bastawade, 1983
L. kharpadi Bastawade, 1986
L. krali Kovarik, 1995
L. lourencoi Kovarik, 1997
L. marmoreus (C.L. Koch, 1844)
L. mjobergi Kraepelin, 1916
L. mucronatus (Fabricius, 1798)
L. nigristernis (Pocock, 1899)
L. obsti Kraepelin, 1913
L. perfidus (Keyserling, 1885)
L. rackae Kovarik, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. variatus (Thorell, 1876)
L. variatus variatus (Thorell, 1876)
L. variatus canopensis Lourenco & Qi, 2007
Lychasioides Vachon, 1974 (1)
L. amieti Vachon, 1974
Mauritanobuthus Qi & Lourenço, 2007* (1)
M. geniezi Qi &Lourenço, 2007*
Mesobuthus Vachon, 1950 (12)
M. afghanus (Pocock, 1889)*
M. agnetis (Werner, 1936)
M. bogdoensis (Birula, 1896)*
M. eupeus (C.L. Koch, 1839)
M. haarlovi Vachon, 1958*
M. iranus (Birula, 1917)*
M. macmahoni (Pocock, 1900)
M. mongolicus (Birula, 1911)*
M. persicus (Pocock, 1899)*
M. phillipsi (Pocock, 1889)*
M. thersites (C. L. Koch, 1839)*
M. vesiculatus (Pocock, 1899)
Mesotityus Gonzalez-Sponga, 1981 (1)
M. vondangeli Gonzalez-Sponga, 1981
Microananteris Lourenço, 2003* (1)
M. minor Lourenço, 2003*
Microbuthus Kraepelin, 1898 (7)
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. gardneri Lowe, 2010*
M. kristensenorum Lowe, 2010*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
M. satyrus Lowe, Kovarik, Stockmann & Stahlavsky, 2018*
Microtityus Kjellesvig-Waering, 1966 (40)
M. angelaerrosae Gonzales-Sponga, 2001*
M. barahona Armas & Teruel, 2012*
M. biordi Gonzalez-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. borincanus Teruel, Rivera & Sanchez, 2014*
M. capayaensis Gonzalez-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae Gonzalez-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. eustatia Armas, 2018*
M. farleyi Teruel, 2000*
M. flavescens Teruel, 2001*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M. jaumei Armas, 1974
M. joseantonioi Gonzalez-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis Gonzalez-Sponga, 2001*
M. lourencoi Armas & Teruel, 2012*
M. minimus Kovarik & Teruel, 2014*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. prendinii Armas & Teruel, 2012*
M. pusillus Teruel & Kovarik, 2012*
M. reini Armas & Teruel, 2012*
M. rickyi Kjellesvig-Waering, 1966
M. santosi Teruel, Rivera & Sanchez, 2014*
M. sevciki Gonzalez-Sponga, 2001*
M. solegladi Armas & Teruel, 2012*
M. starri Lourenço & Huber, 1999*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. vieques Teruel, Rivera & Santos, 2015*
M. virginiae Armas, 1999*
M. vulcanicus Teruel, 2019*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus Gonzalez-Sponga, 2001*
Neobuthus Hirst, 1911 (16)
N. amoudensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. awashensis Kovarik & Lowe, 2012*
N. berberensis Hirst, 1911
N. cloudsleythompsoni Lourenço, 2001*
N. erigavoensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. eritreaensis Lowe & Kovarik, 2016*
N. factorio Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. ferrugineus (Kraepelin, 1898)*
N. gubanensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. haeckeli Kovarik, 2019*
N. kloppersi Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. kutcheri Lowe & Kovarik, 2016*
N. maidensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. montanus Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. solegladi Kovarik, 2019*
N. sudanensis Lourenço, 2005*
Neogrosphus Lourenço, 1995 (3)
N. andrafiabe Lourenço, Wilme & Waeber 2015*
N. blanci Lourenço, 1996
N. griveaudi (Vachon, 1969)
Odontobuthus Vachon, 1950 (6)
O. bidentatus Lourenço & Pezier, 2002*
O. brevidigitus Lowe, 2010*
O. doriae (Thorell, 1876)
O. odonturus (Pocock, 1897)
O. tavighiae Navidpour, Soleglad, Fet & Kovarik, 2013*
O. tirgari Mirshamsi, Azghadi, Navidpour, Aliabadian & Kovarik, 2013*
Odonturus Karsch, 1879 (1)
O. dentatus Karsch, 1879
Olivierus Farzanpay, 1987 (18)
O. bolensis (Sun, Zhu & Lourenço, 2010)*
O. brutus (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. caucasicus (Nordmann, 1840)
O. elenae (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. extremus (Werner, 1936)
O. fuscus (Birula, 1897)*
O. gorelovi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. hainanensis (Birula, 1904)
O. intermedius (Birula, 1897)*
O. karshius (Sun & Sun, 2011)*
O. kaznakovi (Birula, 1904)*
O. kreuzbergi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. longichelus (Sun & Zhu, 2010)*
O. martensii (Karsch, 1879)
O. mischi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. nenilini (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. parthorum (Pocock, 1889)*
O. przewalskii (Birula, 1897)
Orthochiroides Kovarik, 1998 (3)
O. insularis (Pocock, 1899)*
O. socotrensis Kovarik, 2004*
O. vachoni Kovarik, 1998
Orthochirus Karsch, 1891 (53)
O. afar Kovarik, Lowe & Stahlavsky, 2016*
O. afghanus Kovarik, 2004*
O. atarensis Lourenço & Leguin, 2011*
O. bastawadei Zambre, Mirza, Sanap, Upadhye & Javed, 2011*
O. bicolor (Pocock, 1897)
O. birulai Kovarik, Fet & Yagmur, 2020*
O. carinatus Navidpour, Kovarik, Soleglad & Fet, 2019*
O. cloudsleythompsoni Lourenço & Leguin, 2011*
O. danielleae (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fomichevi Kovarik, Yagmur, Fet & Hussen, 2019*
O. formozovi Kovarik, Fet & Yagmur, 2020*
O. fuscipes (Pocock, 1900)
O. gantenbeini Kovarik, Yagmur, Fet & Hussen, 2019*
O. glabrifrons (Kraepelin, 1903)*
O. gromovi Kovarik, 2004*
O. grosseri Kovarik, Fet & Yagmur, 2020*
O. gruberi Kovarik & Fet, 2006*
O. hormozganensis Kovarik & Navidpour, 2020*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. kaspareki (Lourenço & Huber, 2000)*
O. kermanensis Kovarik & Navidpour, 2020*
O. kinzelbachi (Lourenço & Huber, 2000)*
O. krishnai Tikader & Bastawade, 1983
O. kryzhanovskyi Kovarik, Fet & Yagmur, 2020*
O. kucerai Kovarik & Navidpour, 2020*
O. masihipouri Kovarik & Navidpour, 2020*
O. maroccanus Lourenço & Leguin, 2011*
O. melanurus (Kessler, 1874)*
O. mesopotamicus Birula, 1918*
O. minor Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. monodi (Lourenço & Vachon, 1997)*
O. navidpouri Kovarik, Yagmur, Fet & Hussen, 2019*
O. nordmanni Kovarik, Fet & Yagmur, 2020*
O. olivaceus (Karsch, 1881)
O. pallidus (Pocock, 1897)
O. persa (Birula, 1900)*
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. scrobiculosus scrobiculosus (Grube, 1873)
O. scrobiculosus negebensis (Shulov & Amitai, 1960)
O. sejnai Kovarik, Fet & Yagmur, 2020*
O. semnanensis Kovarik & Navidpour, 2020*
O. stockwelli (Lourenço & Vachon, 1995)*
O. tassili Lourenço & Leguin, 2011*
O. tibesti Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. varius Kovarik, 2004*
O. vignolii Kovarik & Navidpour, 2020*
O. zagrosensis Kovarik, 2004*
Pantobuthus Lourenço & Duhem, 2009* (1)
P. complicatus Lourenço & Duhem, 2009*
Parabuthus Pocock, 1890 (38)
P. abyssinicus Pocock, 1901*
P. brevimanus (Thorell, 1876)
P. calvus Purcell, 1898
P. capensis (Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral, 1980
P. erigavoensis Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. eritreaensis Kovarik, 2003*
P. glabrimanus Prendini & Esposito, 2010*
P. gracilis Lamoral, 1979
P. granimanus Pocock, 1895
P. granulatus (Ehrenberg, 1831)
P. hamar Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. heterurus Pocock, 1899
P. hunteri Pocock, 1895
P. kabateki Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. kajibu Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. kalaharicus Lamoral, 1977
P. kraepelini Werner, 1902
P. kuanyamarum Monard, 1937
P. laevifrons (Simon, 1888)
P. liosoma (Ehrenberg, 1828)
P. maximus Werner, 1913
P. mazuchi Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. mossambicensis (Peters, 1861)
P. muelleri Prendini, 2000*
P. namibensis Lamoral, 1979
P. nanus Lamoral, 1979
P. pallidus Pocock, 1895
P. planicauda (Pocock, 1889)
P. raudus (Simon, 1888)
P. robustus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. schlechteri Purcell, 1899
P. setiventer Prendini & Esposito, 2010*
P. somalilandus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. stridulus Hewitt, 1913
P. transvaalicus Purcell, 1899
P. villosus (Peters, 1862)
Pectinibuthus Fet in Orlov & Vasilyev, 1984 (1)
P. birulai Fet in Orlov & Vasilyev, 1984
Physoctonus Mello-Leitao, 1934* (3)
P. amazonicus Lourenço, 2017*
P. debilis (C. L. Koch, 1840)*
P. striatus Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
Picobuthus Lowe, 2010* (2)
P. dundoni Lowe, 2010*
P. wahibaensis Lowe, 2010*
Plesiobuthus Pocock, 1900 (1)
P. paradoxus Pocock, 1900
Polisius Fet, Capes & Sissom, 2001* (1)
P. persicus Fet, Capes & Sissom, 2001*
Pseudolissothus Lourenço, 2001* (1)
P. pusillus Lourenço, 2001*
Pseudolychas Kraepelin, 1911 (3)
P. ochraceus (Hirst, 1911)
P. pegleri (Purcell, 1901)
P. transvaalicus Lawrence, 1961
Pseudouroplectes Lourenço, 1995 (5)
P. betschi Lourenço, 1995
P. lalyae Lourenço & Ythier, 2010*
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999*
P. tsingy Lourenço, Wilme & Waeber, 2016*
Razianus Farzanpay, 1987 (4)
R. birulai Tahir, Navidpour & Prendini, 2014*
R. farzanpayi Tahir, Navidpour & Prendini, 2014*
R. xinjianganus Lourenco, Sun & Zhu, 2010*
R. zarudnyi (Birula, 1903)
Reddyanus Vachon, 1972* (34)
R. aareyensis (Mirza & Sanap, 2010)*
R. acanthurus (Pocock, 1899)*
R. assamensis (Oates, 1888)*
R. basilicus (Karsch, 1879)*
R. besucheti (Vachon, 1982)*
R. bilyi (Kovarik, 2003)*
R. brachycentrus (Pocock, 1899)*
R. ceylonensis Kovarik, Lowe, Ranawana, Hoferek, Jayarathne, Pliskova & Stahlavsky, 2016*
R. corbeti (Tikander & Batawade, 1983)*
R. deharvengi (Lourenço & Duhem, 2010)*
R. feti (Kovarik, 2013)*
R. furai Kovarik & Stahlavsky, 2019*
R. heimi (Vachon, 1976)*
R. hofereki Kovarik & Stahlavsky, 2019*
R. jayarathnei Kovarik, 2016*
R. jendeki (Kovarik, 2013)*
R. justi Kovarik, Lowe & Stahlavsky, 2020*
R. khammamensis (Kovarik, 2003)*
R. krasenskyi (Kovarik, 1998)*
R. kurkai (Kovarik, 1997)*
R. loebli (Vachon, 1982)*
R. majkusi Kovarik & Stahlavsky, 2019*
R. melanodactylus (L. Koch, 1867)*
R. navaiae (Kovarik, 1998)*
R. neradi (Kovarik, 2013)*
R. petrzelkai (Kovarik, 2003)*
R. problematicus (Kovarik, 2003)*
R. ranawanai Kovarik, 2016*
R. rigidulus (Pocock, 1897)*
R. rolciki Kovarik & Stahlavsky, 2019*
R. schwotti Kovarik & Stahlavsky, 2019*
R. tibetanus (Lourenço & Zhu, 2008)*
R. vittatus (Pocock, 1900)
R. zideki (Kovarik, 1994)*
Rhopalurus Thorell, 1876 (3)
R. caribensis Teruel & Roncallo, 2008*
R. laticauda Thorell, 1876
R. ochoai Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
Saharobuthus Lourenço & Duhem, 2009* (1)
S. elegans Lourenço & Duhem, 2009*
Sassanidotus Farzanpay, 1987 (2)
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900)
Somalibuthus Kovarik, 1998 (1)
S. demisi Kovarik, 1998
Somalicharmus Kovarik, 1998 (1)
S. whitmanae Kovarik, 1998
Spelaeolychas Kovarik, 2019* (1)
S. hosei (Pocock, 1891)
Thaicharmus Kovarik, 1995 (4)
T. guptai Mirza, Sanap & Kunte, 2016*
T. indicus Kovarik, 2013*
T. lowei Kovarik, Soleglad & Fet, 2007*
T. mahunkai Kovarik, 1995
Tityobuthus Pocock, 1893 (19)
T. antsingy Lourenço & Goodman, 2004*
T. baroni (Pocock, 1890)
T. betschi Lourenço, Qi & Goodman, 2008*
T. chelbergorum Lourenço, Qi & Goodman, 2008*
T. darainensis Lourenço & Goodman, 2002*
T. dastychi Lourenço, 1997
T. griswoldi Lourenço, 2000*
T. guillaumeti Lourenço, 1995
T. ivohibe Lourenço & Goodman, 1999*
T. judsoni Lourenço, 1996
T. lokobe Lourenço, Waeber & Wilme, 2016*
T. manonae Lourenço, 2000*
T. mccarteri Lourenço, Qi & Goodman, 2008*
T. monodi Lourenço, 2000*
T. pallidus Lourenço, 2004*
T. parrilloi Lourenço, 1996
T. petrae Lourenço, 1996
T. pococki Lourenço, 1995
T. rakotondravonyi Lourenço & Goodman, 2003*
Tityopsis Armas, 1974 (6)
T. canizaresorum Teruel & Rodriguez-Cabrera, 2020*
T. inaequalis (Armas, 1974)
T. inexpectata (Moreno, 1940)
T. mulata Teruel & Rodriguez-Cabrera, 2020*
T. pumila Teruel & Rodriguez-Cabrera, 2020*
T. sheylae Teruel & Rodriguez-Cabrera, 2020*
Tityus C. L. Koch, 1836 (224)
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. abudi Armas, 1999*
T. acananensis Gonzalez-Sponga, 2009*
T. adisi Lourenço & Pezier, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi Gonzalez-Sponga, 2001*
T. alejandroi Teruel, Rivera & Santos, 2015*
T. altithronus Armas, 1999*
T. androcottoides (Karsch, 1879)
T. anduzei Gonzalez-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. anori Lourenço, Rossi & Wilme, 2019*
T. antioquensis Lourenço& Otero Patiño, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai Gonzalez-Sponga, 1985
T. argentinus Borelli, 1899
T. asthenes Pocock, 1893
T. atriventer Pocock, 1897
T. bahiensis (Perty, 1833)
T. bahiensis bahiensis (Perty, 1833)
T. bahiensis eickstedtae Lourenço, 1982
T. barquisimetanus Gonzalez-Sponga, 1994
T. bastosi Lourenço, 1984
T. bellulus Armas, 1999*
T. birabeni Abalos, 1955
T. blanci Lourenço, 1994
T. blaseri Mello-Leitão, 1931
T. boconoensis Gonzalez-Sponga, 1981
T. bolivianus Kraepelin, 1895
T. brazilae Lourenço & Eickstedt, 1984
T. breweri Gonzalez-Sponga, 1997
T. cachipalensis Gonzalez-Sponga, 2002*
T. caesarbarrioi Gonzalez-Sponga, 2001*
T. canopensis Lourenço & Pezier, 2002*
T. carabobensis Gonzalez-Sponga, 1987
T. carinatoides Mello-Leitão, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carolineae Kovarik, Teruel, Cozijn & Seiter, 2013
T. carvalhoi Mello-Leitão, 1945
T. cerroazul Lourenço, 1986
T. championi Pocock, 1898*
T. charalaensis Mello-Leitão, 1940 [nomen nudum]
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. choco Lourenço & Florez D, 2018*
T. cisandinus Lourenço & Ythier, 2017*
T. clathratus C. L. Koch, 1844
T. columbianus (Thorell, 1876)
T. confluens Borelli, 1899
T. confluens confluens Borelli, 1899
T. confluens bodoquena Lourenço, Cabral & Bruehmueller Ramos, 2004
T. costatus (Karsch, 1879)
T. crassicauda Lourenço & Ythier, 2013*
T. crassimanus (Thorell, 1876)
T. cuellari Lourenço, 1994
T. culebrensis Gonzalez-Sponga, 1994
T. curupi Ojanguren-Affilastro, Adilardi, Cajade, Ramirez, Ceccarelli & Mola, 2017*
T. cylindricus (Karsch, 1879)*
T. dasyurus Pocock, 1897
T. dasyurus dasyurus Pocock, 1897
T. dasyurus fulvipes Mello-Leitão, 1945
T. dedoslargos Francke & Stockwell, 1987
T. demangei Lourenço, 1981
T. dillerorum Kovarik, Teruel, Lowe & Friedrich, 2015*
T. dinizi Lourenço, 1997
T. discrepans (Karsch, 1879)
T. dorae Gonzalez-Sponga, 2001*
T. duacaensis Gonzalez-Sponga, 2007*
T. dulceae Gonzalez-Sponga, 2006*
T. dupouyi Gonzalez-Sponga, 1987
T. ecuadorensis Kraepelin, 1896
T. elii Armas & Marcano Fondeur, 1992*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi Gonzalez-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leitão, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessôa, 1935
T. festae Borelli, 1899
T. filodendron Gonzalez-Sponga, 1981
T. florezi Lourenço, 2000*
T. footei Chamberlin, 1916
T. forcipula (Gervais, 1843)
T. fuehrmanni Kraepelin, 1914
T. funestus Hirst, 1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. generaltheophiloi Lourenço, 2017*
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. grahami Lourenço, 2012*
T. grottoedensis Botero-Trujillo & Florez D, 2014*
T. guane Moreno-Gonzalez, Gonzalez O & Florez D, 2019*
T. guaricoensis Gonzalez-Sponga, 2004*
T. haetianus Teruel & Santos, 2018*
T. horacioi Lourenço& Leguin, 2011*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus Mello-Leitão, 1934
T. insignis (Pocock, 1889)
T. intermedius Borelli, 1899
T. isabelceciliae Gonzalez-Sponga, D'Suze & Sevcik, 2001*
T. ivani Gonzalez-Sponga, 2008*
T. ivicnancor Gonzalez-Sponga, 1997
T. jaimei Miranda, Bermudez, Florez D & de Armas, 2020*
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai Gonzalez-Sponga, 2007*
T. karaja Lourenço, 2016*
T. kindli Kovarik & Teruel, 2014*
T. kukututee Ythier, Chevalier & Gangadin, 2020*
T. kuryi Lourenço, 1997
T. lancinii Gonzalez-Sponga, 1972
T. lokiae Lourenço, Adis & Araujo, 2005*
T. longidigitus Gonzalez-Sponga, 2008*
T. lourencoi Flórez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis Gonzalez-Sponga, 2007*
T. mana Lourenço, 2012*
T. manakai Gonzalez-Sponga, 2004*
T. maniapurensis Gonzalez-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. marechali Lourenço, 2013*
T. martinpaechi Lourenço, 2001*
T. matthieseni Lourenço & Pinto-da-Rocha, 2000
T. mattogrossensis Borelli, 1901
T. maturinensis Gonzalez-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis Gonzalez-Sponga, 1974
T. mongei Lourenço, 1996
T. mucusunamensis Gonzalez-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leitão, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leitão, 1940
T. neoespartanus Gonzalez-Sponga, 1996
T. nororientalis Gonzalez-Sponga, 1996
T. obispoi Gonzalez-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus Gonzalez-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis Gonzalez-Sponga, 2007*
T. panguana Kovarik, Teruel, Lowe & Friedrich, 2015*
T. paraguayensis Kraepelin, 1895
T. parvulus Kraepelin, 1914
T. paulistorum Lourenço & da Silva, 2006*
T. perijanensis Gonzalez-Sponga, 1994
T. pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri Gonzalez-Sponga, 1981
T. pococki Hirst, 1907
T. portoplatensis Armas & Marcano Fondeur, 1992
T. potameis Lourenço & Leão Giupponi, 2004*
T. prancei Lourenço, 2000*
T. proseni Abalos, 1954
T. pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis Gonzalez-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus Armas, 1982*
T. ramirezi Esquivel de Verde, 1969 [nomen dubium]
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis Gonzalez-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi Gonzalez-Sponga, 1996
T. romeroi Gonzalez-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rosenbergi Pocock, 1898*
T. rufofuscus Pocock, 1897
T. rupestre Lourenço 2019*
T. rugosus Schenkel, 1932
T. rusmelyae Gonzalez-Sponga, D'Suze & Sevcik, 2001*
T. sabineae Lourenço, 1994
T. sanarensis Gonzalez-Sponga, 1997
T. sanfernandoi Gonzalez-Sponga, 2008*
T. sarisarinamensis Gonzalez-Sponga, 2002*
T. sastrei Lourenço & Flórez, 1990
T. schrammi Teruel & Santos, 2018*
T. serrulatus Lutz & Mello, 1922
T. shiriana Gonzalez-Sponga, 1991
T. silvestris Pocock, 1897
T. simonsi Pocock, 1900
T. smithii Pocock, 1893
T. soratensis Kraepelin, 1911
T. stigmurus (Thorell, 1876)
T. strandi Werner, 1939
T. surimeridensis Gonzalez-Sponga, 2002*
T. surorientalis Gonzalez-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi Gonzalez-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. timendus Pocock, 1898
T. trinitatis Pocock, 1897
T. trivittatus Kraepelin, 1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leitão, 1931
T. unus Lourenço & Pinto-da-Rocha, 2000
T. uquirensis Gonzalez-Sponga, 2001*
T. urachichensis Gonzalez-Sponga, 2007*
T. urbinai Scorza, 1952
T. uruguayensis Borelli, 1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis Gonzalez-Sponga, 1981
T. ventuarensis Gonzalez-Sponga, 2009*
T. wachteli Kovarik, Teruel, Lowe & Friedrich, 2015*
T. walli Gonzalez-Sponga & Wall Gonzalez, 2007*
T. yerenai Gonzalez-Sponga, 2009*
T. ythieri Lourenço, 2007*
T. zulianus Gonzalez-Sponga, 1981
Troglorhopalurus Lourenço, Baptista & Giupponi, 2004* (2)
T. lacrau (Lourenço & Pinto-da-Rocha, 1997)*
T. translucidus Lourenço, Baptista & Giupponi, 2004*
Troglotityobuthus Lourenço, 2000* (1)
T. gracilis (Fage, 1946)
Trypanothacus Lowe, Kovarik, Stockmann & Stahlavsky, 2019* (2)
T. barnesi Lowe, Kovarik, Stockmann & Stahlavsky, 2019*
T. buettikeri (Hendrixson, 2006)*
Uroplectes Peters, 1861 (34)
U. ansiedippenaarae Prendini, 2015*
U. carinatus (Pocock, 1890)
U. carinatus carinatus (Pocock, 1890)
U. carinatus australis Hewitt, 1918
U. chubbi Hirst, 1911
U. emiliae (Werner, 1916)
U. fischeri (Karsch, 1879)
U. flavoviridis Peters, 1861
U. formosus Pocock, 1890
U. formosus formosus Pocock, 1890
U. formosus basuticus Hewitt, 1927
U. formosus maculipes Hewitt, 1918
U. formosus spenceri Pocock, 1896
U. gracilior Hewitt, 1913
U. insignis Pocock, 1890
U. katangensis Prendini, 2015*
U. lineatus (C. L. Koch, 1844)
U. longimanus Werner, 1936
U. machadoi Lourenço, 2000*
U. malawicus Prendini, 2015*
U. marlothi Purcell, 1901
U. ngangelarum Monard, 1930
U. occidentalis Simon, 1876
U. occidentalis occidentalis Simon, 1876
U. occidentalis monardi Vachon, 1950
U. olivaceus Pocock, 1896
U. otjimbinguensis (Karsch, 1879)
U. otjimbinguensis otjimbinguensis (Karsch, 1879)
U. otjimbinguensis massacarum Monard, 1937
U. pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner, 1913
U. pilosus (Thorell, 1876)
U. planimanus (Karsch, 1879)
U. planimanus planimanus (Karsch, 1879)
U. planimanus kuanyamarus Monard, 1937
U. schlechteri Purcell, 1901
U. schubotzi Kraepelin, 1929
U. silvestrii Borelli, 1913
U. teretipes Lawrence, 1966
U. triangulifer (Thorell, 1876)
U. tumidimanus Lamoral, 1979
U. variegatus (C. L. Koch, 1844)
U. vittatus (Thorell, 1876)
U. xanthogrammus Pocock, 1897
U. zambezicus Prendini, 2015*
Vachoniolus Levy, Amitai & Shulov, 1973 (4)
V. batinahensis Lowe, 2010*
V. gallagheri Lowe, 2010*
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
Vachonus Tikader & Bastawade, 1983 (4)
V. asiyaae Amir & Kamaluddin, 2009* [nomen dubium]
V. inexpectatus Lourenço, 2015*
V. iqbali Amir & Kamaluddin, 2009* [nomen dubium]
V. rajasthanicus Tikader & Bastawade, 1983
Xenobuthus Lowe, 2018* (3)
X. anthracinus (Pocock, 1895)*
X. arabicus (Lourenço & Qi, 2006)*
X. xanthus Lowe, 2018*
Zabius Thorell, 1893 (3)
Z. birabeni Mello-Laeitão, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*

Jan Ove Rein (C) 2021