| With 96 genera (one extinct and not listed below) and 1225 species [Updated 22.01.21],
this family is the largest of the scorpion families. This
family is widespread around the world (not found in Antarctica
and New Zealand), and are found in tropical, subtropical and
partly in temperate habitats.
Volschenk, Mattoni & Prendini (2008) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus and Neoprotobuthus) and 15 species are all transfered to Buthidae. August 2019: The family Microcharmidae has been resurrected by Lourenco et al., 2019 and belinging taxa have been transferred back from Buthidae Information about the family Microcharmidae.
Kovarik (2007) has published a revision of the genus Hottentotta. Note that
the medical important Mesobuthus tamulus and two other Asian Mesobuthus have been
transfered to Hottentotta. Kovarik's revision can be
downloaded HERE (browse down to issue 58).
Paraorthochirus has been synonymized with Orthochirus by Navidpour et al., 2008).
The members of this family are small and medium sized
scorpions with a triangular sternum (some genera have a more
pentagonal sternum). In many genera (e.g., Androctonus,
Apistobuthus, Parabuthus, and other Old World
genera) the cauda is usually strong and powerful and the
pedipalps are often very slender whereas other genera have
elongate appendages, especially the males (e.g.,
Centruroides, Lychas, Isometrus and
related genera). Many species are yellow or brown (or
variations of these colors), but black forms are also present.
Some species have quite intricate and attractive patterns
(e.g., Lychas and Isometrus) and colours (e.g.,
Centruroides and Uroplectes). Sizes range from
about 20 mm (like Microtityus and Microbuthus)
to over 120 mm (as in Androctonus, some
Centruroides, Apistobuthus and others).
An indentification key to the genus Compsobuthus was published by Kovarik (2012) [Free full text].
Several species in this family are highly toxic, but fewer
than 20 can be lethal to man. The most notorious species are
found in the genus Androctonus, Centruroides, Hottentotta,
Leiurus, Parabuthus and Tityus. See Scorpions
of medical importance page for further information.
Unlike most other scorpion families, the Buthidae are of
both scientific and public interest due to their medical
importance in many parts of the world. Several species have
also made their way into the hobby. Many of these are safe to
be kept by experienced collectors, but the most dangerous
species should be avoided.
Androctonus crassicauda habitat picture from Kuwait are located HERE.
Photo of Parabuthus leiosoma (left)
by Jan Ove Rein (C)
Photo of Hottentotta sp.
(right) by Jan Ove Rein (C)
SPECIES
FILES:
This list of genera and
species is based on Fet
et al.(2000). September 2019: Subspecies are included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000) |
|
Aegaeobuthus Kovarik, 2019* (4)
A. cyprius (Gantenbein & Kropf, 2000)*
A. gallianoi (Ythier, 2018)*
A. gibbosus (Brulle, 1832)
A. nigrocinctus (Ehrenberg, 1828)*
A. nigrocinctus bishri (Lourenco, 2020)*
|
Afrolychas Kovarik, 2019* (2)
A. braueri (Kraepelin, 1896)
A. burdoi (Simon, 1882)
|
Afroisometrus
Kovarik, 1997 (1)
A. minshullae (Fitzpatrick,
1994)
|
Akentrobuthus
Lamoral, 1976 (2)
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976 |
Alayotityus Armas,
1973 (8)
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A.
juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus
Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas,
1973
|
Ananteris Thorell,
1891 (92)
A. arcadioi Botero-Trujillo, 2008*
A. ashaninka Kovarik, Teruel, Lowe & Friedrich, 2015*
A. ashmolei Lourenço, 1981
A. asuncionensis González-Sponga, 2006*
A. balzanii Thorell, 1891
A. barinensis González-Sponga, 2006*
A. bernabei Giupponi, Vasconcelos & Lourenço, 2009*
A. bianchinii Lourenço, Aguiar-Neto & Limeira-de-Oliveira, 2009*
A. bonito Lourenço, 2012*
A. cachimboensis Lourenço, Motta & da Silva, 2006*
A. camacan Lourenço, Giupponi & Leguin 2013*
A. canalera Miranda & Armas, 2020*
A. capayaensis González-Sponga, 2006*
A. caracensis González-Sponga, 2006*
A. carrasco Lourenço & Motta, 2019*
A. catuaroi González-Sponga, 2006*
A. caucaguitensis González-Sponga, 2006*
A. chagasi Giupponi, Vasconcelos & Lourenço, 2009*
A. charlescorfieldi Lourenço, 2001*
A. chirimakei González-Sponga, 2006*
A. cisandinus Lourenço, 2015*
A. claviformis González-Sponga, 2006*
A. coineaui Lourenço, 1982
A. columbiana Lourenço, 1991
A. cryptozoicus Lourenço, 2005*
A. cumbensis González-Sponga, 2006*
A. curariensis González-Sponga, 2006*
A. cussinii Borelli, 1910
A. dacostai Ythier, Chevalier & Lourenço, 2020*
A. dekeyseri Lourenço, 1982
A. deniseae Lourenço, 1997
A. desiderio Lourenço, Giupponi & Leguin 2013*
A. diegorojasi Rojas-Runjaic, 2005*
A. dorae Botero-Trujillo, 2008*
A. ehrlichi Lourenço, 1994
A. elguapoi González-Sponga, 2006*
A. elisabethae Lourenço, 2003*
A. evellynae Lourenço, 2004*
A. faguasi Botero-Trujillo, 2009*
A. festae Borelli, 1899
A. franckei Lourenço, 1982
A. gorgonae Lourenço & Flórez, 1989
A.
guiripaensis González-Sponga, 2006*
A. guyanensis Lourenço & Monod, 1999*
A. infuscata Lourenço, Giupponi & Leguin 2013*
A. inoae González-Sponga, 2006*
A. intermedia Lourenço, 2012*
A. kalina Ythier, 2018*
A. kuryi Giupponi, Vasconcelos & Lourenço, 2009*
A. leilae Lourenço, 1999*
A. luciae Lourenço, 1984
A. madeirensis Lourenço & Duhem, 2010*
A. mamilihpan Ythier, Chevalier & Lourenço, 2020*
A. maniapurensis González-Sponga, 2006*
A. maranhensis Lourenço, 1987
A. mariaelenae Lourenço, 1999*
A. mariaterezae Lourenço, 1982
A. mauryi Lourenço, 1982
A. meridana González-Sponga, 2006*
A. michaelae Lourenço, 2013*
A. myriamae Botero-Trujillo, 2007*
A. nairae Lourenço, 2004*
A. norae González-Sponga, 2006*
A. ochoai Botero-Trujillo & Florez D, 2011*
A. otavianoi Lira, Pordeus & Ribeiro de Albuquerque, 2017
A. palmari Botero-Trujillo & Noriega, 2011*
A. paoensis González-Sponga, 2006*
A. paracotoensis González-Sponga, 2006*
A. pierrekondre Lourenço, Chevalier, Gangadin & Ythier, 2020*
A. plataensis González-Sponga, 2006*
A. platnicki Lourenço, 1993
A. polleti Lourenço, 2016*
A. principalis González-Sponga, 2006*
A. pydanieli Lourenço, 1982
A. riocaurensis González-Sponga, 2006*
A. riochicoi González-Sponga, 2006*
A. riomachensis Rojas-Runjaic, Portillo-Quintero & Borges, 2008*
A. roraima Lourenço & Duhem, 2010*
A. sabineae Lourenço, 2001*
A. sanchezi González-Sponga, 2006*
A. sepulvedai González-Sponga, 2006*
A. singularis González-Sponga, 2006*
A. sipilili Ythier, Chevalier & Lourenço, 2020*
A. solimariae Botero-Trujillo & Florez D, 2011*
A. surinamensis Lourenço, 2012*
A. tresor Ythier, Chevalier & Lourenço, 2020*
A. terueli Kovarik, 2006*
A. tolimana Teruel & Garcia, 2007*
A. turumbanensis Gonzáles-Sponga, 1980
A.
venezuelensis Gonzáles-Sponga, 1972
A. volschenki Botero-Trujillo, 2009*
A.
zuliana González-Sponga, 2006*
|
Ananteroides
Borelli, 1911* (2)
A. feae Borelli, 1911*
A. inexpectatus Lourenço, 2013*
| |
Androctonus
Ehrenberg, 1828 (29)
A. aeneas C. L. Koch, 1839*
A. afghanus Lourenço & Qi, 2006*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi (Audouin,1826)
A. amoreuxi amoreuxi (Audouin,1826)
A. amoreuxi levyi Fet, 1997
A. australis (Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. barbouri (Werner, 1932)*
A. bicolor Ehrenberg, 1828
A. bicolor bicolor Ehrenberg, 1828
A. bicolor longecarinatus (Caporiacco, 1932) [nomen dubium]
A. cholistanus Kovarik & Ahmed, 2013*
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. donairei Rossi, 2015*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Ythier & Leguin, 2009*
A. pallidus Lourenço, Duhem & Cloudsley-Thompson, 2012*
A. robustus Kovarik & Ahmed, 2013*
A. santi Lourenço, 2015*
A. sergenti Vachon, 1948
A. simonettai Rossi, 2015*
A. tenuissimus Teruel, Kovarik & Turiel, 2013*
A. tigrai Lourenço, Rossi & Sadine 2015*
A. togolensis Lourenço, 2008*
A. tropeai Rossi, 2015*
|
Anomalobuthus Kraepelin, 1900 (6)
A. krivochatskyi Teruel, Kovarik & Fet, 2018*
A. lowei Teruel, Kovarik & Fet, 2018*
A. pavlovskyi Teruel, Kovarik & Fet, 2018*
A. rickmersi Kraepelin, 1900
A. talebii Teruel, Kovarik, Navidpour & Fet, 2014*
A. zarudnyi (Birula, 1911)*
|
Apistobuthus
Finnegan, 1932 (2)
A. pterygocercus Finnegan,
1932
A. susanae Lourenço, 1998
|
Australobuthus
Locket, 1990 (1)
A. xerolimniorum Locket,
1990
|
Babycurus Karsch,
1886 (14)
B. ansorgei Hirst, 1911
B. brignolii Lourenço & Rossi, 2017* [nomen dubium]
B. buettneri Karsch, 1886
B. buettneri buettneri Karsch, 1886
B. buettneri savanicola Lourenco, Bruehmueller Ramos & Cloudsley-Thompson, 2005
B. centrurimorphus Karsch, 1886*
B. dunlopi Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. pictus Pocock, 1896*
B. solegladi Lourenço, 2005*
B. taramassoi Borelli, 1919*
B. wituensis Kraepelin, 1913
|
Baloorthochirus
Kovarik, 1996 (1)
B. becvari Kovarik,
1996
|
Barbaracurus Kovarik, Lowe & Stahlavsky, 2018 (10)
B. feti Kovarik, Lowe &Stahlavsky & Hurre, 2019*
B. exquisitus (Lowe, 2000)*
B. prudenti (Lourenço, 2013)*
B. somalicus (Hirst, 1907)*
B. sofomarensis (Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015)*
B. subpunctatus (Borelli, 1925)*
B. ugartei (Kovarik, 2000)*
B. winklerorum Kovarik, Lowe &Stahlavsky, 2018*
B. yemenensis Kovarik, Lowe &Stahlavsky, 2015*
B. zambonellii (Borelli, 1902)*
| |
Birulatus Vachon,
1974 (3)
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
|
Buthacus Birula,
1908 (23)
B. agarwali Zambre & Lourenço, 2010*
B. ahaggar Lourenço, Kourim & Sadine, 2017*
B. arenicola (Simon, 1885)
B. arenicola arenicola (Simon, 1885)
B. arenicola maroccanus Lourenco, 2006
B. birulai Lourenço, 2006*
B. clevai Lourenço, 2001*
B. elmenia Lourenço & Sadine, 2017*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (nomen dubium)
B. golovatchi Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. leptochelys (Ehrenberg, 1829)
B. leptochelys leptochelys (Ehrenberg, 1829)
B. leptochelys granosus Borelli, 1929
B. leptochelys nitzani Levy, Amitai & Shulov, 1973
B. macrocentrus (Ehrenberg, 1828)*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. occidentalis Vachon, 1953*
B. pakistanensis Lourenço & Qi, 2006*
B. samiae Lourenço & Sadine, 2015*
B. spinatus Lourenço, Bissati & Sadine, 2016*
B. stockmanni Kovarik, Lowe & Stahlavsky, 2016*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. williamsi Lourenço & Leguin, 2009*
B. ziegleri Lourenço, 2000*
|
Butheoloides Hirst, 1925 (19)
B. annieae Lourenço, 1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. granulatus Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. grosseri Kovarik, 2016*
B. hirsti Lourenço, 1996
B. littoralis Lourenço, Touloun & Boumezzough, 2011*
B. maroccanus Hirst, 1925
B. milloti Vachon, 1948
B. monodi Vachon, 1950
B. nuer Kovarik, 2015*
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. savanicola Lourenço, 2013*
B. schwendingeri Lourenço, 2002*
B. slimanii Lourenço, 2010*
B. vanderberghi Lourenço, 2015*
B. wilsoni Lourenço, 1995
|
Butheolus Simon,
1882 (6)
B. andersoni (Pocock, 1895)*
B. gallagheri Vachon, 1980
B. hallani Lourenço & Rossi, 2017*
B. harrisoni Lowe, 2018*
B. thalassinus Simon, 1882
B. villosus Hendrixson, 2006*
|
Buthiscus Birula,
1905 (1)
B. bicalcaratus Birula, 1905
|
Buthoscorpio Werner, 1936 (5)
B. chinnarensis Aswathi, Sureshan & Lourenço, 2015*
B. indicus Lourenço, 2012*
B. politus (Pocock, 1899)
B. rayalensis Javed, Rao, Mirza, Sanap & Tampal, 2010*
B. sarasinorum (Karsch, 1891)
NB! B. jinnahii Amir, Kamaluddin & Jabbar, 2005* and B. rahmatii Amir, Kamaluddin & Jabbar, 2005* was described in the non-valid genus Stenochirus by the authors. I have placed them in Buthoscorpio, which is
the valid name for Stenochirus, but formally this has to be done in a scientific journal before it is valid. Javed et al., 2010 have studied the descriptions of these two species and concluded that they do not belong to the genus Buthoscorpio. This decsision was also supported by Lourenco, 2012. The two taxa are now Buthidae incertae sedis and are not included in the total number species in Buthidae. |
Buthus Leach,
1815 (65)
B. adrianae Rossi, 2013
B. alacanti Teruel & Turiel 2020*
B. albengai Lourenço, 2003*
B. amri Lourenço, Yagmur & Duhem 2010*
B. apiatus Lourenço, El Bouhissi & Sadine, 2020*
B. atlantis Pocock, 1889
B. aures Lourenço & Sadine 2016*
B. awashensis Kovarik, 2011*
B. baeticus Teruel & Turiel 2020*
B. barcaeus Birula, 1909*
B. berberensis Pocock, 1900*
B. bonito Lourenço & Geniez, 2005*
B. boumalenii Touloun & Boumezzough, 2011*
B. boussaadi Lourenço, Chichi & Sadine, 2018*
B. brignolii Lourenço, 2003*
B. centroafricanus Lourenço, 2016*
B. chambiensis Kovarik, 2006*
B. confluens Lourenço, Touloun & Boumezzough, 2012*
B. danyii Rossi, 2017*
B. delafuentei Teruel & Turiel 2020*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. duprei Rossi & Tropea, 2016*
B. egyptiensis Lourenço, 2012*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. elongatus Rossi, 2012*
B. garcialorcai Teruel & Turiel 2020*
B. hassanini Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. ibericus Lourenço & Vachon, 2004*
B. intermedius (Ehrenberg, 1829)*
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. israelis Shulov & Amitai, 1959*
B. jianxinae Lourenço, 2005*
B. karoraensis Rossi & Tropea, 2016*
B. kunti Yagmur, Koc & Lourenço, 2011*
B. labuschagnei Lourenço, 2015*
B. lienhardi Lourenço, 2003*
B. lourencoi Rossi, Tropea & Yagmur, 2013*
B. malhommei Vachon, 1949*
B. manchego Teruel & Turiel 2020*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. nigrovesiculosus Hirst, 1925*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. orientalis Lourenço & Simon, 2012*
B. oudjanii Lourenço, 2017*
B. paris (C. L. Koch, 1839)*
B. parroti Vachon, 1949*
B. pococki Kovarik, Stahlavsky & Elmi, 2020*
B. prudenti Lourenço & Leguin, 2012*
B. pusillus Lourenço, 2013*
B. rochati Lourenço, 2003*
B. saharicus Sadine, Bissati & Lourenço, 2015*
B. serrano Teruel & Turiel 2020*
B. somalilandus Kovarik, Stahlavsky & Elmi, 2020*
B. tassili Lourenço, 2002*
B. trinacrius Lourenço & Rossi, 2013*+
+Probably extinct.
B. tunetanus (Herbst, 1800)*
B. yemenensis Lourenço, 2008*
B. zeylensis Pocock, 1900*
|
Centruroides Marx, 1890 (93)
C. alayoni Armas, 1999*
C. altagraciae Teruel, de Armas & Kovarik, 2015*
C. anchorellus Armas, 1976
C. arctimanus Armas, 1976
C. baergi Hoffmann, 1932*
C. balsasensis Ponce & Francke, 2004*
C. bani
Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis (Pocock, 1898)
C. bertholdii (Thorell, 1876)
C. bicolor (Pocock, 1898)
C. bonito Quijano-Ravell, Teruel & Ponce-Saavedra, 2016*
C. caral Armas & Trujillo, 2013
C. caribbeanus Teurel & Myers, 2017*
C. chamela Ponce-Saavedra & Francke, 2011*
C. chamulaensis Hoffmann, 1932
C. chiapanensis Hoffmann, 1932
C. edwardsii (Gervais, 1843)*
C. elegans (Thorell, 1876)
C. elegans elegans (Thorell, 1876)
C. elegans edentulus Werner, 1939
C. elegans insularis Pocock, 1902
C. exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul (Meise, 1933)
C. fallassisimus Armas & Trujillo, 2010*
C. farri Armas, 1976
C. flavopictus (Pocock, 1898)
C. flavopictus flavopictus (Pocock, 1898)
C. flavopictus meridionalis Hoffmann, 1932
C. franckei Santibanez-Lopez & Contreras-Felix, 2013*
C. fulvipes (Pocock, 1898)
C. galano Teruel, 2001*
C. gracilis (Latreille, 1804)
C. granosus (Thorell, 1876)*
C. griseus (C. L. Koch, 1844)
C. guanensis Franganillo, 1930
C. hentzi (Banks, 1900)
C. hirsuticauda Teruel, 2011*
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. huichol Teruel, Ponce-Saavedra & Quijano-Ravell, 2015*
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. ixil Trujillo & Armas, 2016*
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. lauriadnae Ponce-Saavedra & Francke, 2019*
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. lucidus Teruel, Armas & Kovarik, 2015*
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mascota Ponce-Saavedra & Francke, 2011*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus (Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. panamensis Arias & Esposito, 2014*
C. platnicki Armas, 1981
C. pococki Sissom & Francke, 1983
C. polito Teruel, 2007*
C. poncei Teruel, Kovarik, Baldazo-Monsivais & Hoferek, 2015*
C. possanii Gonzalez-Santillan, Galan-Sanchez & Valdez-Velazquez, 2019*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. rodolfoi Santibanez-Lopez & Contreras-Felix, 2013*
C. romeroi Quijano-Ravell, De Armas, Francke & Ponce-Saavedra, 2019*
C. ruana Quijano-Ravell & Ponce-Saavedra, 2016*
C. sanandres Armas, Sarmiento & Florez 2012*
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. simplex (Thorell, 1876)*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. suffusus (Pocock, 1902)
C. suffusus suffusus (Pocock, 1902)
C. tapachulaensis Hoffmann, 1932*
C. tecomanus Hoffmann, 1932*
C. tenuis Thorell, 1876*
C. testaceus (DeGeer, 1778)
C. thorellii (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. villegasi Baldazo-Monsivaiz, Ponce-Saavedra & Flores-Moreno, 2013*
C. vittatus (Say, 1821)
|
Charmus Karsch,
1879 (5)
C. brignolii Lourenço, 2000*
C. indicus Hirst, 1915
C. laneus Karsch, 1879
C. saradieli Kovarik, Lowe, Ranawana, Hoferek & Jayarathne 2016*
C. sinhagadensis Tikader & Bastawade, 1983
|
Chaneke Francke, Teruel & Santibanez-Lopez, 2014* (4)
C. aliciae (Armas & Frias, 1998)*
C. baldazoi Kovarik, Teruel & Lowe, 2016*
C. fogoso Francke, Teruel & Santibanez-Lopez, 2014*
C. hofereki Kovarik, Teruel & Lowe, 2016*
|
Cicileiurus Teruel,
2007* (1)
C. monticola Teruel, 2007*
|
Cicileus Vachon,
1948 (5)
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
C. hoggarensis Lourenço & Rossi, 2015*
C. latellai Lourenço & Rossi, 2015*
C. montanus Lourenço & Rossi, 2015*
|
Compsobuthus Vachon, 1949 (49)
C. abyssinicus (Birula, 1903)
C. acutecarinatus (Simon, 1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov, 1973
C. armenicus Lourenço, Leguin & Duhem, 2015*
C. atrostriatus (Pocock, 1897)*
C. berlandi Vachon, 1950
C. birulai Lourenço, Leguin & Duhem, 2010*
C. boucheti Lourenço, Duhem & Cloudsley-Thompson, 2012*
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. eritreaensis Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. garyi Lourenço & Vachon, 2001*
C. humaae Amir, Kamaluddin & Kahn, 2005* (nomen dubium)
C. jakesi Kovarik, 2003*
C. jordanensis
Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. klaptoczi (Birula, 1909)
C. krali Kovarik, 2012*
C. levyi Kovarik, 2012*
C. longipalpis Levy, Amitai & Shulov, 1973
C. maidensis Kovarik, 2018*
C. maindroni (Kraepelin, 1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. plutenkoi Kovarik, 2003*
C. polisi Lowe, 2001*
C. rugosulus (Pocock,
1900)
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. sindicus Kovarik & Ahmed, 2011*
C. somalilandus Kovarik, 2012*
C. tassili Lourenço, 2010*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. turieli Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. ullrichi Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. vachoni Sisson, 1994
C. werneri (Birula, 1908)
|
Congobuthus Lourenço,
1999* (1)
C. fagei Lourenço, 1999*
|
Darchenia Vachon,
1977 (1)
D. bernadettae Vachon, 1977
|
Egyptobuthus Lourenço,
1999* (1)
E. vaissadei Lourenço, 1999*
|
Femtobuthus Lowe, 2010* (1)
F. shutuae Lowe, 2010*
|
Fetilinia Lowe & Kovarik, 2021* (1)
F. dentator Lowe & Kovarik, 2021*
|
|
Gint Kovarik, Lowe, Pliskova & Stahlavsky, 2013* (12)
G. amoudensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. banfasae Kovarik & Lowe, 2019*
G. calviceps (Pocock, 1900)
G. childsi Kovarik, 2018*
G. dabakalo Kovarik & Mazuch, 2015*
G. gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013*
G. gubanensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. insolitus (Borelli, 1925)* (nomen dubium)
G. maidensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. marialuisae Rossi, 2015* (nomen dubium)
G. monicae Rossi, 2015* (nomen dubium)
G. puntlandus Kovarik & Mazuch, 2015*
|
Grosphus Simon,
1880 (34)
G. ambre Lourenco, Wilmé & Waeber, 2018*
G. ankarafantsika Lourenço, 2003*
G. ankarana Lourenço & Goodman, 2003*
G. annulatus Fage, 1929*
G. bemaraha Lourenco, Wilmé & Waeber, 2018*
G. bicolor Lourenço, 2012*
G. bistriatus Kraepelin, 1900*
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. eliseanneae Lourenço & Wilme, 2016*
G. feti Lourenço, 1996*
G. flavopiceus Kraepelin, 1900*
G. ganzhorni Lourenço, Wilme & Waeber, 2016*
G. grandidieri Kraepelin, 1900*
G. goudoti Lourenço & Goodman, 2006*
G. halleuxi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. hirtus Kraepelin, 1900
G. intertidalis Lourenço, 1999*
G. limbatus (Pocock, 1889)*
G. madagascariensis (Gervais, 1843)
G. magalieae Lourenço, 2014*
G. makay Lourenço & Wilme, 2015*
G. mandena Lourenço, 2005*
G. mahafaliensis Lourenço, Goodman & Ramilijaona, 2004*
G. mavo Lourenço & Rossi 2020*
G. mayottensis Lourenço & Goodman, 2009*
G. olgae Lourenço, 2004*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. rakotoariveloi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. rossii Lourenço, 2013*
G. sabineae Lourenço & Wilme, 2016*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
G. tavaratra Lourenco, Soarimalala & Goodman, 2009*
G. voahangyae Lourenço & Wilme 2015*
G. waeberi Lourenço & Wilme, 2016*
|
Hemibuthus Pocock, 1900 (2)
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
|
Hemilychas Hirst,
1911 (1)
H. alexandrinus (Hirst, 1911)
|
Heteroctenus Pocock, 1893* (9)
H. abudi (Armas & Marcano Fondeur, 1987)*
H. aridicola (Teruel et Armas, 2012)*
H. bonettii (Armas 1999)*
H. garridoi (Armas, 1974)*
H. gibarae (Teruel, 2006)*
H. granulimanus (Teruel, 2006)*
H. junceus (Herbst, 1800)*
H. melloleitaoi (Teruel et Armas, 2006)*
H. princeps (Karsch, 1879)*
|
Himalayotityobuthus
Lourenço, 1997 (2)
H. alejandrae Lourenço, 2003*
H. martensi Lourenço,
1997
| |
Hottentotta
Birula, 1908 (55)
H. alticola (Pocock, 1895)
H. alticola alticola (Pocock, 1895)
H. alticola nigrifrons (Pocock, 1900)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004* (nomen dubium)
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. flavidulus Teruel & Rein, 2010*
H. franzwerneri (Birula, 1914)
H. fuscitruncus (Caporiacco, 1936)
H. gambelaensis Kovarik, 2015*
H. gentili (Pallary, 1924)*
H. gibaensis Kovarik, 2015*
H. hoggarensis Lourenço & Leguin, 2014*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. juliae Kovarik, Yagmur & Fet, 2019*
H. keralaensis Aswhati, Sureshan & Lourenço, 2016*
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. krivokhatskyi Kovarik, Yagmur & Fet, 2019*
H. lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarik & Kayedi, 2010*
H. mazuchi Kovarik, 2013*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1875)
H. minax minax (L. Koch, 1975)
H. minax occidentalis (Vachon & Stockmann, 1968)
H. minusalta Vachon, 1959*
H. navidpouri Kovarik, Yagmur & Moradi, 2018*
H. niloticus (Birula, 1928)*
H. novaki Kovarik, 2015*
H. pachyurus (Pocock, 1897)*
H. pellucidus Lowe, 2010*
H. penjabensis (Birula, 1897)*
H. polystictus (Pocock, 1896)
H. reddyi Lourenço, 2015*
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1880)
H. saxinatans Lowe, 2010*
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. sistanensis Kovarik, Yagmur & Moradi, 2018*
H. socotrensis (Pocock, 1889)
H. somalicus Kovarik, 2018*
H. songi (Lourenço, Qi & Zhu, 2005)*
H. sousai Turiel, 2014*
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trailini Kovarik, 2013*
H. trilineatus (Peters, 1861)
H. ugandaensis Kovarik, 2013*
H. vinchu Mirza, Ambekar & Kulkarni, 2019*
H. zagrosensis Kovarik, 1997
|
Iranobuthus
Kovarik, 1997 (1)
I. krali Kovarik,
1997
|
Ischnotelson Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (2)
I. guanambiensis (Lenarducci, Pinto-da-Rocha & Lucas, 2005)*
I. peruassu Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
|
Isometroides
Keyserling, 1885 (1)
I. vescus (Karsch,
1880)
|
Isometrus Ehrenberg,
1828 (10)
I. amboli Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. atherii Amir & Kamaluddin, 2008*
I. formosus Pocock, 1894
I. isadensis Tikader & Bastawade, 1983*
I. kovariki Sulakhe, Dandekar, Mukherjee, Pandey, Ketkar, Padhye & Bastawade, 2020*
I. liaqatii Amir & Kamaluddin, 2008*
I. maculatus (DeGeer, 1778)
I. tamhini Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. thurstoni Pocock, 1893
I. thwaitesi Pocock, 1897
|
Jaguajir Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (3)
J. agamemnon (C. L. Koch, 1839)*
J. pintoi (Mello-Leitão, 1932)*
J. pintoi pintoi (Mello-Leitão, 1932)*
J. pintoi kourouensis (Lourenco, 2008)*
J. rochae (Borelli, 1910)*
|
Janalychas Kovarik, 2019* (9)
J. albimanus (Henderson, 1919)
J. farkasi (Kovarik, 1997)
J. granulatus Mirza, 2020*
J. heurtaultae (Kovarik, 1997)
J. keralaensis Mirza, 2020*
J. laevifrons (Pocock, 1897)
J. shoplandi (Oates, 1888)
J. srilankensis (Lorenco, 1997)
J. tricarinatus (Simon, 1884)
|
Karasbergia
Hewitt, 1913 (1)
K. methueni Hewitt, 1913 |
Kraepelinia
Vachon, 1974 (1)
K. palpator (Birula, 1903) |
Lanzatus Kovarik,
2001* (2)
L. somalicus Kovarik, 2001*
L. somalilandus Kovarik, Lowe & Stahlavsky, 2016* |
Leiurus Ehrenberg, 1828 (17)
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. arabicus Lowe, Yagmur & Kovarik, 2014*
L. ater Lourenço, 2019*
L. brachycentrus (Ehrenberg, 1829)*
L. dekeyseri Lourenço, 2020*
L. gubanensis Kovarik & Lowe, 2020*
L. haenggii Lowe, Yagmur & Kovarik, 2014*
L. heberti Lowe, Yagmur & Kovarik, 2014*
L. hebraeus (Birula, 1908)*
L. hoggarensis Lourenço, Kourim & Sadine, 2018*
L. kuwaiti Lourenço, 2020*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. macroctenus Lowe, Yagmur & Kovarik, 2014*
L. quinquestriatus (Ehrenberg, 1828)
L. saharicus Lourenço, 2020*
L. savanicola Lourenço, Qi & Cloudsley-Thompson, 2006*
L. somalicus Lourenço, & Rossi, 2016*
| |
Liobuthus Birula,
1898 (1)
L. kessleri Birula, 1898 |
Lissothus Vachon, 1948 (3)
L. bernardi Vachon, 1948
L. chaambi Lourenço & Sadine, 2014*
L. occidentalis Vachon, 1950 |
Lychas C.L. Koch,
1845 (31)
L. aberlenci Lourenço, 2013*
L. armasi Kovarik, 2013*
L. armillatus (Gervais, 1841)*
L. asper (Pocock, 1891)
L. biharensis Tikader & Bastawade, 1983
L. brehieri Lourenço, 2017*
L. buchardi Kovarik, 1997
L. cernickai Kovarik, 2013*
L. flavimanus (Thorell,
1888)
L. gravelyi Henderson, 1913
L. hendersoni (Pocock, 1897)
L. hillyardi Kovarik, 1997
L. inexpectatus Lourenço, 2011*
L. kamshetensis Tikader & Bastawade, 1983
L. kharpadi Bastawade, 1986
L. krali Kovarik, 1995
L. Lourençoi Kovarik, 1997
L. marmoreus (C.L. Koch, 1844)
L. mjobergi Kraepelin, 1916
L. mucronatus (Fabricius, 1798)
L. nigristernis (Pocock, 1899)
L. obsti Kraepelin, 1913
L. perfidus (Keyserling, 1885)
L. rackae Kovarik, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. variatus (Thorell, 1876)
L. variatus variatus (Thorell, 1876)
L. variatus canopensis Lourenco & Qi, 2007
|
|
Lychasioides
Vachon, 1974 (1)
L. amieti Vachon, 1974 |
Mauritanobuthus Qi & Lourenço, 2007* (1)
M. geniezi Qi &Lourenço, 2007*
|
Mesobuthus Vachon, 1950 (12)
M. afghanus (Pocock, 1889)*
M. agnetis (Werner, 1936)
M. bogdoensis (Birula, 1896)*
M. eupeus (C.L. Koch, 1839)
M. haarlovi Vachon, 1958*
M. iranus (Birula, 1917)*
M. macmahoni (Pocock, 1900)
M. mongolicus (Birula, 1911)*
M. persicus (Pocock, 1899)*
M. phillipsi (Pocock, 1889)*
M. thersites (C. L. Koch, 1839)*
M. vesiculatus (Pocock,
1899)
|
Mesotityus
Gonzalez-Sponga, 1981 (1)
M. vondangeli
González-Sponga, 1981 |
Microananteris
Lourenço, 2003* (1)
M. minor Lourenço, 2003* |
|
Microbuthus
Kraepelin, 1898 (7)
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. gardneri Lowe, 2010*
M. kristensenorum Lowe, 2010*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
M. satyrus Lowe, Kovarik, Stockmann & Stahlavsky, 2018*
|
|
Microtityus
Kjellesvig-Waering, 1966 (40)
M. angelaerrosae Gonzáles-Sponga, 2001*
M. barahona Armas & Teruel, 2012*
M. biordi González-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. borincanus Teruel, Rivera & Sanchez, 2014*
M. capayaensis González-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae González-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. eustatia Armas, 2018*
M. farleyi Teruel, 2000*
M. flavescens Teruel, 2001*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M. jaumei Armas, 1974
M. joseantonioi González-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis González-Sponga, 2001*
M. lourencoi Armas & Teruel, 2012*
M. minimus Kovarik & Teruel, 2014*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. prendini Armas & Teruel, 2012*
M. pusillus Teruel & Kovarik, 2012*
M. reini Armas & Teruel, 2012*
M. rickyi Kjellesvig-Waering, 1966
M. santosi Teruel, Rivera & Sanchez, 2014*
M. sevciki González-Sponga, 2001*
M. solegladi Armas & Teruel, 2012*
M. starri Lourenço & Huber, 1999*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. vieques Teruel, Rivera & Santos, 2015*
M. virginiae Armas, 1999*
M. vulcanicus Teruel, 2019*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus González-Sponga, 2001*
|
Neobuthus Hirst,
1911 (16)
N. amoudensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. awashensis Kovarik & Lowe, 2012*
N. berberensis (Hirst, 1911)
N. cloudsleythompsoni Lourenço, 2001*
N. erigavoensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. eritreaensis Lowe & Kovarik, 2016*
N. factorio Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. ferrugineus (Kraepelin, 1898)*
N. gubanensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. haeckeli Kovarik, 2019*
N. kloppersi Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. kutcheri Lowe & Kovarik, 2016*
N. maidensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. montanus Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. solegladi Kovarik, 2019*
N. sudanensis Lourenço, 2005*
|
Neogrosphus Lourenço, 1995 (3)
N. andrafiabe Lourenço, Wilme & Waeber 2015*
N. blanci Lourenço, 1996
N. griveaudi (Vachon, 1969) |
Odontobuthus Vachon, 1950 (6)
O. bidentatus (Lourenço & Pezier, 2002)*
O. brevidigitus Lowe, 2010*
O. doriae (Thorell, 1876)
O. odonturus (Pocock, 1897)
O. tavighiae Navidpour, Soleglad, Fet & Kovarik, 2013*
O. tirgari Mirshamsi, Azghadi, Navidpour, Aliabadian & Kovarik, 2013*
|
Odonturus Karsch,
1879 (1)
O.
dentatus Karsch, 1879 |
Olivierus Farzanpay, 1987 (18)
O. bolensis (Sun, Zhu & Lourenço, 2010)*
O. brutus (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. caucasicus (Nordmann, 1840)
O. elenae (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. extremus (Werner, 1936)
O. fuscus (Birula, 1897)*
O. gorelovi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. hainanensis (Birula, 1904)
O. intermedius (Birula, 1897)*
O. karshius (Sun & Sun, 2011)*
O. kaznakovi (Birula, 1904)*
O. kreuzbergi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. longichelus (Sun & Zhu, 2010)*
O. martensii (Karsch, 1879)
O. mischi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. nenilini (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. parthorum (Pocock, 1889)*
O. przewalskii (Birula, 1897)
|
-
Orthochiroides
Kovarik, 1998 (3)
O. insularis (Pocock, 1889)*
O. socotrensis Kovarik, 2004*
O. vachoni Kovarik, 1998
|
Orthochirus
Karsch, 1891 (52)
O. afar Kovarik, Lowe & Stahlavsky, 2016*
O. afghanus Kovarik, 2004*
O. atarensis Lourenço & Leguin, 2011*
O. bastawadei Zambre, Mirza, Sanap, Upadhye & Javed, 2011*
O. bicolor (Pocock, 1897)
O. bicolor bicolor (Pocock, 1897)
O. bicolor insularis (Pocock, 1899)
O. birulai Kovarik, Fet & Yagmur, 2020*
O. carinatus Navidpour, Kovarik, Soleglad & Fet, 2019*
O. cloudsleythompsoni Lourenço & Leguin, 2011*
O. danielleae (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fomichevi Kovarik, Yagmur, Fet & Hussen, 2019*
O. formozovi Kovarik, Fet & Yagmur, 2020*
O. fuscipes (Pocock, 1900)
O. gantenbeini Kovarik, Yagmur, Fet & Hussen, 2019*
O. glabrifrons (Kraepelin, 1903)*
O. gromovi Kovarik, 2004*
O. grosseri Kovarik, Fet & Yagmur, 2020*
O. gruberi Kovarik & Fet, 2006*
O. hormozganensis Kovarik & Navidpour, 2020*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. kaspareki (Lourenço & Huber, 2000)*
O. kermanensis Kovarik & Navidpour, 2020*
O. kinzelbachi (Lourenço & Huber, 2000)*
O. krishnai Tikader & Bastawade, 1983
O. kryzhanovskyi Kovarik, Fet & Yagmur, 2020*
O. kucerai Kovarik & Navidpour, 2020*
O. masihipouri Kovarik & Navidpour, 2020*
O. maroccanus Lourenço & Leguin, 2011*
O. mesopotamicus Birula, 1918*
O. minor Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. monodi (Lourenço & Vachon, 1997)*
O. navidpouri Kovarik, Yagmur, Fet & Hussen, 2019*
O. nordmanni Kovarik, Fet & Yagmur, 2020*
O. olivaceus (Karsch, 1881)
O. pallidus (Pocock, 1897)
O. persa (Birula, 1900)*
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. scrobiculosus scrobiculosus (Grube, 1873)
O. scrobiculosus negebensis (Shulov & Amitai, 1960)
O. sejnai Kovarik, Fet & Yagmur, 2020*
O. semnanensis Kovarik & Navidpour, 2020*
O. stockwelli (Lourenço & Vachon, 1995)*
O. tassili Lourenço & Leguin, 2011*
O. tibesti Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. varius Kovarik, 2004*
O. vignolii Kovarik & Navidpour, 2020*
O. zagrosensis Kovarik, 2004*
|
Pantobuthus
Lourenço & Duhem, 2009* (1)
P. complicatus Lourenço & Duhem, 2009*
|
Parabuthus Pocock,
1890 (38)
P. abyssinicus (Pocock, 1901)*
P. brevimanus (Thorell, 1876)
P. calvus Purcell, 1898
P. capensis (Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral, 1980
P. erigavoensis Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. eritreaensis Kovarik, 2003*
P. glabrimanus Prendini & Esposito, 2010*
P. gracilis Lamoral, 1979
P. granimanus Pocock, 1895
P. granulatus (Ehrenberg, 1831)
P. hamar Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. heterurus Pocock, 1899
P. hunteri Pocock, 1895
P. kabateki Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. kajibu Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. kalaharicus Lamoral, 1977
P. kraepelini Werner, 1902
P. kuanyamarum Monrad, 1937
P. laevifrons (Simon, 1888)
P. liosoma (Ehrenberg, 1828)
P. maximus Werner, 1913
P. mazuchi Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. mossambicensis (Peters, 1861)
P. muelleri Prendini, 2000*
P. namibensis Lamoral, 1979
P. nanus Lamoral, 1979
P. pallidus Pocock, 1895
P. planicauda (Pocock, 1889)
P. raudus (Simon, 1888)
P. robustus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. schlechteri Purcell, 1899
P. setiventer Prendini & Esposito, 2010*
P. somalilandus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. stridulus Hewitt, 1913
P.
transvaalicus Purcell, 1899
P.
villosus (Peters, 1862)
|
Pectinibuthus Fet in Orlov & Vasilyev, 1984 (1)
P. birulai Fet in Orlov & Vasilyev, 1984 |
Physoctonus
Mello-Leitao, 1934* (3)
P. amazonicus Lourenço, 2017*
P. debilis (C. L. Koch, 1840)*
P. striatus Esposito, Yamaguti, Souza, Pinto da Roacha & Prendini, 2017*
|
Picobuthus Lowe, 2010* (2)
P. dundoni Lowe, 2010*
P. wahibaensis Lowe, 2010*
|
Plesiobuthus
Pocock, 1900 (1)
P. paradoxus Pocock, 1900
|
Polisius Fet,
Capes & Sissom, 2001* (1)
P. persicus Fet,
Capes & Sissom, 2001* |
|
Pseudolissothus
Lourenço, 2001* (1)
P. pussilus Lourenço, 2001*
|
Pseudolychas
Kraepelin, 1911 (3)
P. ochraceus (Hirst,
1911)
P. pegleri (Purcell, 1901)
P.
transvaalicus Lawrence, 1961
|
|
Pseudouroplectes
Lourenço, 1995 (5)
P. betschi Lourenço, 1995
P. lalyae Lourenço & Ythier, 2010*
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999*
P. tsingy Lourenço, Wilme & Waeber, 2016*
|
Razianus Farzanpay, 1987 (4)
R. birulai Tahir, Navidpour & Prendini, 2014*
R. farzanpayi Tahir, Navidpour & Prendini, 2014*
R. xinjianganus Lourenco, Sun & Zhu, 2010*
R. zarudnyi (Birula, 1903)
|
Reddyanus Vachon, 1972* (34)
R. aareyensis (Mirza & Sanap, 2010)*
R. acanthurus (Pocock, 1899)*
R. assamensis (Oates, 1888)*
R. basilicusi (Karsch, 1879)*
R. besucheti (Vachon, 1982)*
R. bilyi (Kovarik, 2003)*
R. brachycentrus (Pocock, 1899)*
R. ceylonensis Kovarik, Lowe, Ranawana, Hoferek, Jayarathne & Stahlavsky, 2016*
R. corbeti (Tikander & Batawade, 1983)*
R. deharvengi (Lourenço & Duhem, 2010)*
R. feti (Kovarik, 2013)*
R. furai Kovarik & Stahlavsky, 2019*
R. heimi (Vachon, 1976)*
R. hofereki Kovarik & Stahlavsky, 2019*
R. jayarathnei Kovarik, 2016*
R. jendeki (Kovarik, 2013)*
R. justi Kovarik, Lowe & Stahlavsky, 2020*
R. khammamensis (Kovarik, 2003)*
R. krasenskyi (Kovarik, 1998)*
R. kurkai (Kovarik, 1997)*
R. loebli (Vachon, 1982)*
R. majkusi Kovarik & Stahlavsky, 2019*
R. melanodactylus (L. Koch, 1867)*
R. navaiae (Kovarik, 1998)*
R. neradi (Kovarik, 2013)*
R. petrzelkai (Kovarik, 2003)*
R. problematicus (Kovarik, 2003)*
R. ranawanai Kovarik, 2016*
R. rigidulus (Pocock, 1897)*
R. rolciki Kovarik & Stahlavsky, 2019*
R. schwotti Kovarik & Stahlavsky, 2019*
R. tibetanus (Lourenço & Zhu, 2008)*
R. vittatus (Pocock, 1900)
R. zideki (Kovarik, 1994)*
|
|
Rhopalurus Thorell, 1876 (3)
R. caribensis Teruel & Roncallo, 2008*
R. laticauda Thorell, 1876
R. ochoai Esposito, Yamaguti, Souza, Pinto da Roacha & Prendini, 2017*
|
|
|
|
Saharobuthus
Lourenço & Duhem, 2009* (1)
S. elegans Lourenço & Duhem, 2009*
|
Sassandiothus
Farzanpay, 1987 (2)
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900)
|
Somalibuthus
Kovarik, 1998 (1)
S. demisi Kovarik, 1998 |
Somalicharmus
Kovarik, 1998 (1)
S. withmanae Kovarik, 1998 |
Spelaeolychas Kovarik, 2019* (1)
S. hosei (Pocock, 1891)
|
Thaicharmus Kovarik, 1995 (4)
T. guptai Mirza, Sanap & Kunte, 2016*
T. indicus Kovarik, 1995
T. lowei Kovarik, Soleglad & Fet, 2007*
T. mahunkai Kovarik, 1995
|
|
Tityobuthus
Pocock, 1893 (19)
T. antsingy Lourenço & Goodman, 2004*
T. baroni (Pocock, 1890)
T. betschi Lourenço, Qi & Goodman, 2008*
T. chelbergorum Lourenço, Qi & Goodman, 2008*
T. darainensis Lourenço & Goodman, 2002*
T. dastychi Lourenço, 1997
T. griswoldi Lourenço, 2000*
T. guillaumeti Lourenço, 1995
T. ivohibe Lourenço & Goodman, 1999*
T. judsoni Lourenço, 1996
T. lokobe Lourenço, Waeber & Wilme, 2016*
T. manonae Lourenço, 2000*
T. mccarteri Lourenço, Qi & Goodman, 2008*
T. monodi Lourenço, 2000*
T. pallidus Lourenço, 2004*
T.
parrilloi Lourenço, 1996
T. petrae
Lourenço, 1996
T. pococki Lourenço, 1995
T. rakotondravonyi Lourenço, 2003*
|
Tityopsis Armas,
1974 (6)
T. canizaresorum Teruel & Rodriguez-Cabrera, 2020*
T. inaequalis (Armas, 1974)
T. inexpectata (Moreno, 1940)
T. mulata Teruel & Rodriguez-Cabrera, 2020*
T. pumila Teruel & Rodriguez-Cabrera, 2020*
T. sheylae Teruel & Rodriguez-Cabrera, 2020*
|
Tityus C. L. Koch,
1836 (223)
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. abudi Armas, 1999*
T. acananensis González-Sponga, 2009*
T. adisi Lourenço, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi González-Sponga, 2001*
T. alejandroi Teruel, Rivera & Santos, 2015*
T. altithronus Armas, 1999*
T. androcottoides (Karsch, 1879)
T. anduzei González-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. antioquensis Lourenço& Otero Patiño, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai González-Sponga, 1985
T. argentinus Borelli, 1899
T. asthenes Pocock, 1893
T. atriventer Pocock, 1897
T. bahiensis (Perty, 1833)
T. bahiensis bahiensis (Perty, 1833)
T. bahiensis eickstedtae Lourenço, 1982
T. barquisimetanus González-Sponga, 1994
T. bastosi Lourenço, 1984
T. bellulus Armas, 1999*
T. birabeni Abalos, 1955
T. blanci Lourenço, 1994
T. blaseri Mello-Leitão, 1931
T. boconoensis González-Sponga, 1981
T. bolivanus Kraepelin, 1895
T. brazilae Lourenço & Eickstedt, 1984
T. breweri González-Sponga, 1997
T. cachipalensis González-Sponga, 2002*
T. caesarbarrioi González-Sponga, 2001*
T. canopensis Lourenço & Pezier, 2002*
T. carabobensis González-Sponga, 1987
T. carinatoides Mello-Leitão, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carolineae Kovarik, Teruel, Cozijn & Seiter, 2013
T. carvalhoi Mello-Leitão, 1945
T. cerroazul Lourenço, 1986
T. championi Pocock, 1898*
T. charalaensis Mello-Leitão, 1940 [nomen nudum]
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. choco Lourenço & Florez D, 2018*
T. cisandinus Lourenço & Ythier, 2017*
T. clathratus C. L. Koch, 1844
T. columbianus (Thorell, 1876)
T. confluens Borelli, 1899
T. confluens confluens Borelli, 1899
T. confluens bodoquena Lourenço, Cabral & Bruehmueller Ramos, 2004
T. costatus (Karsch, 1879)
T. crassicauda Lourenço, 2013*
T. crassimanus (Thorell, 1876)
T. cuellari Lourenço, 1994
T. culebrensis González-Sponga, 1994
T. curupi Ojanguren-Affilastro, Adilardi, Cajade, Ramõarez, Ceccarelli & Mola, 2017*
T. cylindricus (Karsch, 1879)*
T. dasyurus Pocock, 1897
T. dasyurus dasyurus Pocock, 1897
T. dasyurus fulvipes Mello-Leitão, 1945
T. dedoslargos Francke & Stockwell, 1987
T. demangei Lourenço, 1981
T. dillerorum Kovarik, Teruel, Lowe & Friedrich, 2015*
T. dinizi Lourenço, 1997
T. discrepans (Karsch, 1879)
T. dorae González-Sponga, 2001*
T. duacaensis González-Sponga, 2007*
T. dulceae González-Sponga, 2006*
T. dupouyi González-Sponga, 1987
T. ecuadorensis
Kraepelin, 1896
T. elii (Armas & Marcano
Fondeur, 1992)*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi González-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leitão, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessôa, 1935
T. festae Borelli, 1899
T. filodendron González-Sponga, 1981
T. florezi Lourenço, 2000*
T. footei Chamberlin, 1916
T. forcipula (Gervais, 1843)
T. fuhrmanni Kraepelin, 1914
T. funestus Hirst, 1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. generaltheophiloi Lourenço, 2017*
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. grahami Lourenço, 2012*
T. grottoedensis Botero-Trujillo & Florez D, 2014*
T. guane Moreno-Gonzalez, Gonzalez O & Florez D, 2019*
T. guariconesis Gonzalez-Sponga, 2004*
T. haetianus Teruel & Santos, 2018*
T. horacioi Lourenço& Leguin, 2011*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus Mello-Leitão, 1934
T. insignis (Pocock, 1889)
T. intermedius Borelli, 1899
T. isabelceciliae González-Sponga, D'Suze & Sevcik, 2001*
T. ivani González-Sponga, 2008*
T. ivicnancor González-Sponga,
1997
T. jaimei Miranda, Bermudez, Florez D & de Armas, 2020*
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai González-Sponga, 2007*
T. karaja Lourenço, 2016*
T. kindli KOvarik & Teruel, 2014*
T. kukututee Ythier, Chevalier & Gangadin, 2020*
T. kuryi Lourenço, 1997
T. lancinii González-Sponga, 1972
T. lokiae Lourenço, 2005*
T. longidigitus González-Sponga, 2008*
T. Lourençoi Flórez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis González-Sponga, 2007*
T. mana Lourenço, 2012*
T. manakai González-Sponga, 2004*
T. maniapurensis González-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. marechali Lourenço, 2013*
T. martinpaechi Lourenço, 2001*
T. matthieseni Lourenço & Pinto-da-Rocha, 2000
T. mattogrossensis Borelli,
1901
T. maturinensis González-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis González-Sponga, 1974
T. mongei Lourenço, 1996
T. mucusunamensis González-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leitão, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leitão, 1940
T. neoespartanus González-Sponga, 1996
T. nororientalis González-Sponga, 1996
T. obispoi González-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus González-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis González-Sponga, 2007*
T. panguana Kovarik, Teruel, Lowe & Friedrich, 2015*
T. paraguayensis Kraepelin, 1895
T.
parvulus Kraepelin, 1914
T. paulistorum Lourenço & Qi, 2006*
T.
perijanensis González-Sponga, 1994
T.
pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri
González-Sponga, 1981
T. pococki Hirst,
1907
T. portoplatensis Armas & Marcano
Fondeur, 1992
T. potameis Lourenço & Leão Giupponi, 2004*
T. prancei Lourenço,
2000*
T. proseni Abalos, 1954
T.
pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis González-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus (Armas, 1982)*
T. ramirezi Esquivel de Verde, 1969 [nomen dubium]
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis González-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi González-Sponga, 1996
T. romeroi González-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rosenbergi Pocock, 1898*
T. rufofuscus Pocock, 1897
T. rupestre Lourenço 2019*
T. rugosus Schenkel, 1932
T. rusmelyae González-Sponga, D'Suze & Sevcik, 2001*
T. sabinae Lourenço, 1994
T. sanarensis González-Sponga, 1997
T. sanfernandoi González-Sponga, 2008*
T. sarisarinamensis González-Sponga, 2002*
T. sastrei Lourenço & Flórez, 1990
T. schrammi Teruel & Santos, 2018*
T. serrulatus Lutz & Mello, 1922
T. shiriana González-Sponga, 1991
T. silvestris Pocock, 1897
T. simonsi Pocock, 1900
T. smithii Pocock, 1893
T. soratensis Kraepelin, 1912
T. stigmurus (Thorell, 1876)
T. strandi Werner, 1939
T. surimeridensis González-Sponga, 2002*
T. surorientalis González-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi González-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. timendus Pocock, 1898
T. trinitatis Pocock, 1897
T. trivittatus Kraepelin, 1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leitão, 1931
T. unus Lourenço & Pinto-da-Rocha, 2000
T. uquirensis González-Sponga, 2001*
T. urachichensis González-Sponga, 2007*
T. urbinai
Scorza, 1952
T. uruguayensis Borelli,
1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis González-Sponga, 1981
T. ventuarensis González-Sponga, 2009*
T. wachteli Kovarik, Teruel, Lowe & Friedrich, 2015*
T. walli González-Sponga & Wall Gonzalez, 2007*
T. yerenai González-Sponga, 2009*
T. ythieri Lourenço, 2007*
T. zulianus González-Sponga, 1981
|
|
Troglorhopalurus Lourenço, Baptista & Giupponi,
2004* (2)
T. lacrau (Lourenço & Pinto-da-Rocha, 1997)*
T. translucidus Lourenço, Baptista & Giupponi, 2004*
| Troglotityobuthus Lourenço,
2000* (1)
T. gracilis (Fage, 1946)
|
Trypanothacus Lowe, Kovarik, Stockmann & Stahlavsky, 2019* (2)
T. barnesi Lowe, Kovarik, Stockmann & Stahlavsky, 2019*
T. buettikeri (Hendrixson, 2006)*
| |
Uroplectes Peters,
1861 (34)
U. ansiedippenaarae Prendini, 2015*
U. carinatus (Pocock, 1890)
U. carinatus carinatus (Pocock, 1890)
U. carinatus australis Hewitt, 1918
U. chubbi Hirst, 1911
U. emiliae (Werner, 1916)
U. fischeri (Karsch, 1879)
U. flavoviridis Peters, 1861
U. formosus Pocock, 1890
U. formosus formosus Pocock, 1890
U. formosus basuticus Hewitt, 1927
U. formosus maculipes Hewitt, 1918
U. formosus spenceri Pocock, 1896
U. gracilior Hewitt, 1914
U. insignis Pocock, 1890
U. katangensis Prendini, 2015*
U. lineatus (C. L. Koch, 1844)
U. longimanus Werner, 1936
U. machadoi Lourenço, 2000*
U. malawicus Prendini, 2015*
U. marlothi Purcell, 1901
U. ngangelarum Monard, 1930
U. occidentalis Simon, 1876
U. occidentalis occidentalis Simon, 1876
U. occidentalis monardi Vachon, 1950
U. olivaceus Pocock, 1896
U. otjimbinguensis (Karsch, 1879)
U. otjimbinguensis otjimbinguensis (Karsch, 1879)
U. otjimbinguensis massacarum Monard, 1937
U. pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner, 1913
U. pilosus (Thorell, 1876)
U. planimanus (Karsch, 1879)
U. planimanus planimanus (Karsch, 1879)
U. planimanus kuanyamarus Monard, 1937
U. schlechteri Purcell, 1901
U. schubotzi Kraepelin, 1929
U. silvestrii Borelli, 1913
U. teretipes Lawrence, 1966
U. triangulifer (Thorell, 1876)
U. tumidimanus Lamoral, 1979
U. variegatus (C. L. Koch, 1844)
U. vittatus (Thorell, 1876)
U. xanthogrammus Pocock, 1897
U. zambezicus Prendini, 2015*
| |
Vachoniolus Levy, Amitai & Shulov, 1973 (4)
V. batinahensis Lowe, 2010*
V. gallagheri Lowe, 2010*
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
|
Vachonus Tikader & Bastawade, 1983 (4)
V. asiyaae Amir & Kamaluddin, 2009* [nomen dubium]
V. inexpectatus Lourenço, 2015*
V. iqbali Amir & Kamaluddin, 2009* [nomen dubium]
V. rajasthanicus Tikader & Bastawade, 1983
|
Xenobuthus Lowe, 2018* (3)
X. anthracinus (Pocock, 1895)*
X. arabicus (Lourenço & Qi, 2006)*
X. xanthus Lowe, 2018*
|
Zabius Thorell, 1893 (3)
Z. birabeni Mello-Laeitão, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*
|
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