| With 99 genera (one extinct and not listed below) and 1349 species [Updated 03.02.23],
this family is the largest of the scorpion families. This
family is widespread around the world (not found in Antarctica
and New Zealand), and are found in tropical, subtropical and
partly in temperate habitats.
26.07.22: Lowe & Kovarik (2022) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus and Neoprotobuthus) and 17 species are all transfered to Buthidae. August 2019: The family Microcharmidae has been resurrected by Lourenco et al., 2019 and belinging taxa have been transferred back from Buthidae Information about the family Microcharmidae.
Kovarik (2007) has published a revision of the genus Hottentotta. Note that
the medical important Mesobuthus tamulus and two other Asian Mesobuthus have been
transfered to Hottentotta. Kovarik's revision can be
downloaded HERE (browse down to issue 58).
Paraorthochirus has been synonymized with Orthochirus by Navidpour et al., 2008).
The members of this family are small and medium sized
scorpions with a triangular sternum (some genera have a more
pentagonal sternum). In many genera (e.g., Androctonus,
Apistobuthus, Parabuthus, and other Old World
genera) the cauda is usually strong and powerful and the
pedipalps are often very slender whereas other genera have
elongate appendages, especially the males (e.g.,
Centruroides, Lychas, Isometrus and
related genera). Many species are yellow or brown (or
variations of these colors), but black forms are also present.
Some species have quite intricate and attractive patterns
(e.g., Lychas and Isometrus) and colours (e.g.,
Centruroides and Uroplectes). Sizes range from
about 20 mm (like Microtityus and Microbuthus)
to over 120 mm (as in Androctonus, some
Centruroides, Apistobuthus and others).
An indentification key to the genus Compsobuthus was published by Kovarik (2012) [Free full text].
Several species in this family are highly toxic, but fewer
than 20 can be lethal to man. The most notorious species are
found in the genus Androctonus, Centruroides, Hottentotta,
Leiurus, Parabuthus and Tityus. See Scorpions
of medical importance page for further information.
Unlike most other scorpion families, the Buthidae are of
both scientific and public interest due to their medical
importance in many parts of the world. Several species have
also made their way into the hobby. Many of these are safe to
be kept by experienced collectors, but the most dangerous
species should be avoided.
Androctonus crassicauda habitat picture from Kuwait are located HERE.
Photo of Parabuthus leiosoma (left)
by Jan Ove Rein (C)
Photo of Hottentotta sp.
(right) by Jan Ove Rein (C)
SPECIES
FILES:
This list of genera and
species is based on Fet
et al.(2000). September 2019: Subspecies are included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000) |
|
Aegaeobuthus Kovarik, 2019* (4)
A. cyprius (Gantenbein & Kropf, 2000)*
A. gallianoi (Ythier, 2018)*
A. gibbosus (Brulle, 1832)
A. nigrocinctus (Ehrenberg, 1828)*
A. nigrocinctus bishri (Lourenco, 2020)*
|
Afrolychas Kovarik, 2019* (2)
A. braueri (Kraepelin, 1896)
A. burdoi (Simon, 1882)
|
Afroisometrus
Kovarik, 1997 (1)
A. minshullae (Fitzpatrick,
1994)
|
Akentrobuthus
Lamoral, 1976 (2)
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976 |
Alayotityus Armas,
1973 (8)
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A.
juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus
Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas,
1973
|
Ananteris Thorell,
1891 (96)
A. arcadioi Botero-Trujillo, 2008*
A. ashaninka Kovarik, Teruel, Lowe & Friedrich, 2015*
A. ashmolei Lourenço, 1981
A. asuncionensis González-Sponga, 2006*
A. balzanii Thorell, 1891
A. barinensis González-Sponga, 2006*
A. bernabei Giupponi, Vasconcelos & Lourenço, 2009*
A. bianchinii Lourenço, Aguiar-Neto & Limeira-de-Oliveira, 2009*
A. bonito Lourenço, 2012*
A. cachimboensis Lourenço, Motta & da Silva, 2006*
A. camacan Lourenço, Giupponi & Leguin 2013*
A. canalera Miranda & Armas, 2020*
A. capayaensis González-Sponga, 2006*
A. caracensis González-Sponga, 2006*
A. carrasco Lourenço & Motta, 2019*
A. catuaroi González-Sponga, 2006*
A. caucaguitensis González-Sponga, 2006*
A. chagasi Giupponi, Vasconcelos & Lourenço, 2009*
A. charlescorfieldi Lourenço, 2001*
A. chirimakei González-Sponga, 2006*
A. cisandinus Lourenço, 2015*
A. claviformis González-Sponga, 2006*
A. coineaui Lourenço, 1982
A. columbiana Lourenço, 1991
A. cryptozoicus Lourenço, 2005*
A. cumbensis González-Sponga, 2006*
A. curariensis González-Sponga, 2006*
A. cussinii Borelli, 1910
A. dacostai Ythier, Chevalier & Lourenço, 2020*
A. dekeyseri Lourenço, 1982
A. deniseae Lourenço, 1997
A. desiderio Lourenço, Giupponi & Leguin 2013*
A. diegorojasi Rojas-Runjaic, 2005*
A. dorae Botero-Trujillo, 2008*
A. ehrlichi Lourenço, 1994
A. elguapoi González-Sponga, 2006*
A. elisabethae Lourenço, 2003*
A. evellynae Lourenço, 2004*
A. faguasi Botero-Trujillo, 2009*
A. festae Borelli, 1899
A. franckei Lourenço, 1982
A. gorgonae Lourenço & Florez, 1989
A.
guiripaensis González-Sponga, 2006*
A. guyanensis Lourenço & Monod, 1999*
A. infuscata Lourenço, Giupponi & Leguin 2013*
A. inoae González-Sponga, 2006*
A. intermedia Lourenço, 2012*
A. kalina Ythier, 2018*
A. karupina Lourenço, 2021*
A. kayapo Lourenço, 2022*
A. kuryi Giupponi, Vasconcelos & Lourenço, 2009*
A. leilae Lourenço, 1999*
A. luciae Lourenço, 1984
A. madeirensis Lourenço & Duhem, 2010*
A. mamilihpan Ythier, Chevalier & Lourenço, 2020*
A. maniapurensis González-Sponga, 2006*
A. maranhensis Lourenço, 1987
A. mariaelenae Lourenço, 1999*
A. mariaterezae Lourenço, 1982
A. martensi Lourenço, 2021*
A. mauryi Lourenço, 1982
A. meridana González-Sponga, 2006*
A. michaelae Lourenço, 2013*
A. myriamae Botero-Trujillo, 2007*
A. nairae Lourenço, 2004*
A. norae González-Sponga, 2006*
A. obscura Lourenço & Motta, 2021*
A. ochoai Botero-Trujillo & Florez D, 2011*
A. otavianoi Lira, Pordeus & Ribeiro de Albuquerque, 2017
A. palmari Botero-Trujillo & Noriega, 2011*
A. paoensis González-Sponga, 2006*
A. paracotoensis González-Sponga, 2006*
A. pierrekondre Lourenço, Chevalier, Gangadin & Ythier, 2020*
A. plataensis González-Sponga, 2006*
A. platnicki Lourenço, 1993
A. polleti Lourenço, 2016*
A. principalis González-Sponga, 2006*
A. pydanieli Lourenço, 1982
A. riocaurensis González-Sponga, 2006*
A. riochicoi González-Sponga, 2006*
A. riomachensis Rojas-Runjaic, Portillo-Quintero & Borges, 2008*
A. roraima Lourenço & Duhem, 2010*
A. sabineae Lourenço, 2001*
A. sanchezi González-Sponga, 2006*
A. sepulvedai González-Sponga, 2006*
A. singularis González-Sponga, 2006*
A. sipilili Ythier, Chevalier & Lourenço, 2020*
A. solimariae Botero-Trujillo & Florez D, 2011*
A. surinamensis Lourenço, 2012*
A. tresor Ythier, Chevalier & Lourenço, 2020*
A. terueli Kovarik, 2006*
A. tolimana Teruel & Garcia, 2007*
A. turumbanensis Gonzales-Sponga, 1980
A.
venezuelensis Gonzales-Sponga, 1972
A. volschenki Botero-Trujillo, 2009*
A.
zuliana González-Sponga, 2006*
|
Ananteroides
Borelli, 1911* (2)
A. feae Borelli, 1911*
A. inexpectatus Lourenço, 2013*
| |
Androctonus Ehrenberg, 1828 (36)
A. aeneas C. L. Koch, 1839*
A. afghanus Lourenço & Qi, 2006*
A. agrab Ythier & Lourenço, 2022*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi (Audouin,1826)
A. amoreuxi amoreuxi (Audouin,1826)
A. amoreuxi levyi Fet, 1997
A. australis (Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. barbouri (Werner, 1932)*
A. bartolozzii Rossi & Merendino, 2016*
A. bicolor Ehrenberg, 1828
A. bicolor bicolor Ehrenberg, 1828
A. bicolor longecarinatus (Caporiacco, 1932) [nomen dubium]
A. bourdoni Vachon, 1948*
A. burkinensis Ythier, 2021*
A. cholistanus Kovarik & Ahmed, 2013*
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. donairei Rossi, 2015*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Ythier & Leguin, 2009*
A. pallidus Lourenço, Duhem & Cloudsley-Thompson, 2012*
A. robustus Kovarik & Ahmed, 2013*
A. santi Lourenço, 2015*
A. sergenti Vachon, 1948
A. simonettai Rossi, 2015*
A. sistanus Barahoei & Mirshamsi, 2022*
A. tenuissimus Teruel, Kovarik & Turiel, 2013*
A. tibesti Lourenço & El-Hennawy, 2022*
A. tigrai Lourenço, Rossi & Sadine 2015*
A. togolensis Lourenço, 2008*
A. tropeai Rossi, 2015*
A. turkiyensis Yagmur, 2021*
|
Anomalobuthus Kraepelin, 1900 (6)
A. krivochatskyi Teruel, Kovarik & Fet, 2018*
A. lowei Teruel, Kovarik & Fet, 2018*
A. pavlovskyi Teruel, Kovarik & Fet, 2018*
A. rickmersi Kraepelin, 1900
A. talebii Teruel, Kovarik, Navidpour & Fet, 2014*
A. zarudnyi (Birula, 1911)*
|
Apistobuthus
Finnegan, 1932 (2)
A. pterygocercus Finnegan,
1932
A. susanae Lourenço, 1998
|
Australobuthus
Locket, 1990 (1)
A. xerolimniorum Locket,
1990
|
Babycurus Karsch,
1886 (14)
B. ansorgei Hirst, 1911
B. brignolii Lourenço & Rossi, 2017* [nomen dubium]
B. buettneri Karsch, 1886
B. buettneri buettneri Karsch, 1886
B. buettneri savanicola Lourenco, Bruehmueller Ramos & Cloudsley-Thompson, 2005
B. centrurimorphus Karsch, 1886*
B. dunlopi Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. pictus Pocock, 1896*
B. solegladi Lourenço, 2005*
B. taramassoi Borelli, 1919*
B. wituensis Kraepelin, 1913
|
Baloorthochirus
Kovarik, 1996 (1)
B. becvari Kovarik,
1996
|
Barbaracurus Kovarik, Lowe & Stahlavsky, 2018 (11)
B. feti Kovarik, Lowe &Stahlavsky & Hurre, 2019*
B. exquisitus (Lowe, 2000)*
B. kabateki Kovarik, Lowe, Stahlavsky & Just, 2022*
B. prudenti (Lourenço, 2013)*
B. somalicus (Hirst, 1907)*
B. sofomarensis (Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015)*
B. subpunctatus (Borelli, 1925)*
B. ugartei (Kovarik, 2000)*
B. winklerorum Kovarik, Lowe &Stahlavsky, 2018*
B. yemenensis Kovarik, Lowe &Stahlavsky, 2018*
B. zambonellii (Borelli, 1902)*
| |
Birulatus Vachon,
1974 (4)
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
B. jordanensis Lourenço, Al-Saraireh, Afifeh, Baker, Bader-Katbeh & Amr, 2021*
|
Buthacus Birula,
1908 (35)
B. agarwali Zambre & Lourenço, 2010*
B. ahaggar Lourenço, Kourim & Sadine, 2017*
B. algerianus Lourenço, 2006*
B. amitaii Cain, Gefen & Prendini, 2021*
B. arava Cain, Gefen & Prendini, 2021*
B. arenicola (Simon, 1885)
B. armasi Lourenço, 2013*
B. bicolor Afifeh, Al-Saraireh, Baker, Amr & Lourenço, 2022*
B. birulai Lourenço, 2006*
B. clevai Lourenço, 2001*
B. elmenia Lourenço & Sadine, 2017*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (nomen dubium)
B. fuscata Pallary, 1929*
B. golovatchi Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. leptochelys (Ehrenberg, 1829)
B. levyi Cain, Gefen & Prendini, 2021*
B. maroccanus Lourenço, 2006*
B. mahraouii Lourenço, 2004*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. nitzani Levy, Amitai & Shulov, 1973
B. occidentalis Vachon, 1953*
B. pakistanensis Lourenço & Qi, 2006*
B. sadinei Ythier, 2022*
B. samiae Lourenço & Sadine, 2015*
B. spatzi (Birula, 1911)
B. spinatus Lourenço, Bissati & Sadine, 2016*
B. stockmanni Kovarik, Lowe & Stahlavsky, 2016*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. williamsi Lourenço & Leguin, 2009*
B. yotvatensis Levy, Amitai & Shulov, 1973
B. ziegleri Lourenço, 2000*
|
Butheoloides Hirst, 1925 (19)
B. annieae Lourenço, 1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. granulatus Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. grosseri Kovarik, 2016*
B. hirsti Lourenço, 1996
B. littoralis Lourenço, Touloun & Boumezzough, 2011*
B. maroccanus Hirst, 1925
B. milloti Vachon, 1948
B. monodi Vachon, 1950
B. nuer Kovarik, 2015*
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. savanicola Lourenço, 2013*
B. schwendingeri Lourenço, 2002*
B. slimanii Lourenço, 2010*
B. vanderberghi Lourenço, 2015*
B. wilsoni Lourenço, 1995
|
Butheolus Simon,
1882 (6)
B. andersoni (Pocock, 1895)*
B. gallagheri Vachon, 1980
B. hallani Lourenço & Rossi, 2017*
B. harrisoni Lowe, 2018*
B. thalassinus Simon, 1882
B. villosus Hendrixson, 2006*
|
Buthiscus Birula,
1905 (1)
B. bicalcaratus Birula, 1905
|
Buthoscorpio Werner, 1936 (5)
B. chinnarensis Aswathi, Sureshan & Lourenço, 2015*
B. indicus Lourenço, 2012*
B. politus (Pocock, 1899)
B. rayalensis Javed, Rao, Mirza, Sanap & Tampal, 2010*
B. sarasinorum (Karsch, 1891)
NB! B. jinnahii Amir, Kamaluddin & Jabbar, 2005* and B. rahmatii Amir, Kamaluddin & Jabbar, 2005* was described in the non-valid genus Stenochirus by the authors. I have placed them in Buthoscorpio, which is
the valid name for Stenochirus, but formally this has to be done in a scientific journal before it is valid. Javed et al., 2010 have studied the descriptions of these two species and concluded that they do not belong to the genus Buthoscorpio. This decsision was also supported by Lourenco, 2012. The two taxa are now Buthidae incertae sedis and are not included in the total number species in Buthidae. |
Buthus Leach, 1815 (78)
B. adrianae Rossi, 2013
B. ahaggar Ythier, Sadine, Haddadi & Lourenco 2021*
B. ajax (C. L. Koch, 1839)*
B. alacanti Teruel & Turiel 2020*
B. albengai Lourenço, 2003*
B. amri Lourenço, Yagmur & Duhem 2010*
B. apiatus Lourenço, El Bouhissi & Sadine, 2020*
B. atlantis Pocock, 1889
B. aures Lourenço & Sadine 2016*
B. awashensis Kovarik, 2011*
B. baeticus Teruel & Turiel 2020*
B. balmensis Ythier & Laborieux, 2022*
B. barcaeus Birula, 1909*
B. berberensis Pocock, 1900*
B. bobo Ythier, 2021*
B. bonito Lourenço & Geniez, 2005*
B. boumalenii Touloun & Boumezzough, 2011*
B. boussaadi Lourenço, Chichi & Sadine, 2018*
B. brignolii Lourenço, 2003*
B. castellano Teruel & Turiel 2022*
B. centroafricanus Lourenço, 2016*
B. chambiensis Kovarik, 2006*
B. confluens Lourenço, Touloun & Boumezzough, 2012*
B. danyii Rossi, 2017*
B. delafuentei Teruel & Turiel 2020*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. duprei Rossi & Tropea, 2016*
B. egyptiensis Lourenço, 2012*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. elongatus Rossi, 2012*
B. gabani Ythier, 2021*
B. garcialorcai Teruel & Turiel 2020*
B. goyffoni Abidi, Sadine & Lourenco 2021*
B. halius (C. L. Koch, 1839)*
B. hassanini Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. iaspis Teruel & Turiel 2022*
B. ibericus Lourenço & Vachon, 2004*
B. intermedius (Ehrenberg, 1829)*
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. israelis Shulov & Amitai, 1959*
B. jianxinae Lourenço, 2005*
B. karoraensis Rossi & Tropea, 2016*
B. kunti Yagmur, Koc & Lourenço, 2011*
B. labuschagnei Lourenço, 2015*
B. lienhardi Lourenço, 2003*
B. lourencoi Rossi, Tropea & Yagmur, 2013*
B. malhommei Vachon, 1949*
B. manchego Teruel & Turiel 2020*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. nabataeus Lourenço, Abu Afifeh & Al-Saraireh, 2021*
B. nigrovesiculosus Hirst, 1925*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. orientalis Lourenço & Simon, 2012*
B. oudjanii Lourenço, 2017*
B. paris (C. L. Koch, 1839)*
B. parroti Vachon, 1949*
B. pedrosousai Teruel & Turiel 2021*
B. pococki Kovarik, Stahlavsky & Elmi, 2020*
B. prudenti Lourenço & Leguin, 2012*
B. pusillus Lourenço, 2013*
B. pyrenaeus Ythier, 2021*
B. rochati Lourenço, 2003*
B. saharicus Sadine, Bissati & Lourenço, 2015*
B. serrano Teruel & Turiel 2020*
B. somalilandus Kovarik, Stahlavsky & Elmi, 2020*
B. tassili Lourenço, 2002*
B. trinacrius Lourenço & Rossi, 2013*+
+Probably extinct.
B. tunetanus (Herbst, 1800)*
B. turkana Ythier & Lourenço, 2022*
B. yemenensis Lourenço, 2008*
B. zeylensis Pocock, 1900*
|
Centruroides Marx, 1890 (101)
C. alayoni Armas, 1999*
C. altagraciae Teruel, de Armas & Kovarik, 2015*
C. anchorellus Armas, 1976
C. arctimanus Armas, 1976
C. baergi Hoffmann, 1932*
C. baldazoi Ponce-Saavedra, Linares-Guillen & Quijano-Ravell, 2022*
C. balsasensis Ponce & Francke, 2004*
C. bani
Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis (Pocock, 1898)
C. berstoni Goodman, Prendini, Francke & Esposito, 2021*
C. bertholdii (Thorell, 1876)
C. bicolor (Pocock, 1898)
C. bonito Quijano-Ravell, Teruel & Ponce-Saavedra, 2016*
C. caral Armas & Trujillo, 2013
C. caribbeanus Teruel & Myers, 2017*
C. chamela Ponce-Saavedra & Francke, 2011*
C. chamulaensis Hoffmann, 1932
C. catemacoensis Goodman, Prendini, Francke & Esposito, 2021*
C. chanae Goodman, Prendini, Francke & Esposito, 2021*
C. chiapanensis Hoffmann, 1932
C. concordia Armas & Teruel, 2021*
C. cuauhmapan Goodman, Prendini, Francke & Esposito, 2021*
C. edwardsii (Gervais, 1843)*
C. elegans (Thorell, 1876)
C. elegans elegans (Thorell, 1876)
C. elegans edentulus Werner, 1939
C. elegans insularis Pocock, 1902
C. exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul (Meise, 1933)
C. fallassisimus Armas & Trujillo, 2010*
C. farri Armas, 1976
C. flavopictus (Pocock, 1898)
C. flavopictus flavopictus (Pocock, 1898)
C. flavopictus meridionalis Hoffmann, 1932
C. franckei Santibanez-Lopez & Contreras-Felix, 2013*
C. fulvipes (Pocock, 1898)
C. galano Teruel, 2001*
C. gracilis (Latreille, 1804)
C. granosus (Thorell, 1876)*
C. griseus (C. L. Koch, 1844)
C. guanensis Franganillo, 1930
C. hamadryas Goodman, Prendini, Francke & Esposito, 2021*
C. hentzi (Banks, 1900)
C. hirsuticauda Teruel, 2011*
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. huichol Teruel, Ponce-Saavedra & Quijano-Ravell, 2015*
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. ixil Trujillo & Armas, 2016*
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. lauriadnae Ponce-Saavedra & Francke, 2019*
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. lucidus Teruel, Armas & Kovarik, 2015*
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mascota Ponce-Saavedra & Francke, 2011*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus (Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. panamensis Arias & Esposito, 2014*
C. platnicki Armas, 1981
C. pococki Sissom & Francke, 1983
C. polito Teruel, 2007*
C. poncei Teruel, Kovarik, Baldazo-Monsivais & Hoferek, 2015*
C. possanii Gonzalez-Santillan, Galan-Sanchez & Valdez-Velazquez, 2019*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. rodolfoi Santibanez-Lopez & Contreras-Felix, 2013*
C. romeroi Quijano-Ravell, De Armas, Francke & Ponce-Saavedra, 2019*
C. ruana Quijano-Ravell & Ponce-Saavedra, 2016*
C. sanandres Armas, Sarmiento & Florez 2012*
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. simplex (Thorell, 1876)*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. suffusus Pocock, 1902
C. tapachulaensis Hoffmann, 1932*
C. tecomanus Hoffmann, 1932*
C. tenuis Thorell, 1876*
C. testaceus (DeGeer, 1778)
C. thorellii (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. villegasi Baldazo-Monsivaiz, Ponce-Saavedra & Flores-Moreno, 2013*
C. vittatus (Say, 1821)
C. yucatanensis Goodman, Prendini, Francke & Esposito, 2021*
|
Charmus Karsch,
1879 (5)
C. brignolii Lourenço, 2000*
C. indicus Hirst, 1915
C. laneus Karsch, 1879
C. saradieli Kovarik, Lowe, Ranawana, Hoferek & Jayarathne 2016*
C. sinhagadensis Tikader & Bastawade, 1983
|
Chaneke Francke, Teruel & Santibanez-Lopez, 2014* (4)
C. aliciae (Armas & Frias, 1998)*
C. baldazoi Kovarik, Teruel & Lowe, 2016*
C. fogoso Francke, Teruel & Santibanez-Lopez, 2014*
C. hofereki Kovarik, Teruel & Lowe, 2016*
|
Cicileiurus Teruel,
2007* (1)
C. monticola Teruel, 2007*
|
Cicileus Vachon,
1948 (5)
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
C. hoggarensis Lourenço & Rossi, 2015*
C. latellai Lourenço & Rossi, 2015*
C. montanus Lourenço & Rossi, 2015*
|
Compsobuthus Vachon, 1949 (54)
C. abyssinicus (Birula, 1903)
C. air Lourenço & Rossi, 2010*
C. acutecarinatus (Simon, 1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov, 1973
C. armenicus Lourenço & Leguin, 2015*
C. atrostriatus (Pocock, 1897)*
C. berlandi Vachon, 1950
C. birulai Lourenço, Leguin & Duhem, 2010*
C. boucheti Lourenço, Duhem & Cloudsley-Thompson, 2012*
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. eritreaensis Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. garyi Lourenço & Vachon, 2001*
C. humaae Amir, Kamaluddin & Kahn, 2005* (nomen dubium)
C. jakesi Kovarik, 2003*
C. jordanensis
Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. khaybari Abu Afifeh, Aloufi & Al-Saraireh, 2021*
C. klaptoczi (Birula, 1909)
C. krali Kovarik, 2012*
C. levyi Kovarik, 2012*
C. longipalpis Levy, Amitai & Shulov, 1973
C. mahazat Ythier & Lourenço, 2023*
C. maidensis Kovarik, 2018*
C. maindroni (Kraepelin, 1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. plutenkoi Kovarik, 2003*
C. polisi Lowe, 2001*
C. rugosulus (Pocock, 1900)
C. satpuraensis Waghe, Gangalmale & Khandekar, 2022*
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. sindicus Kovarik & Ahmed, 2011*
C. somalilandus Kovarik, 2012*
C. tassili Lourenço, 2010*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. turieli Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. ullrichi Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. vachoni Sisson, 1994
C. vannii Rossi, 2016*
C. werneri (Birula, 1908)
|
Congobuthus Lourenço,
1999* (1)
C. fagei Lourenço, 1999*
|
Darchenia Vachon,
1977 (1)
D. bernadettae Vachon, 1977
|
Egyptobuthus Lourenço,
1999* (1)
E. vaissadei Lourenço, 1999*
|
Femtobuthus Lowe, 2010* (1)
F. shutuae Lowe, 2010*
|
Fetilinia Lowe & Kovarik, 2021* (1)
F. dentator Lowe & Kovarik, 2021*
|
|
Gint Kovarik, Lowe, Pliskova & Stahlavsky, 2013* (12)
G. amoudensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. banfasae Kovarik & Lowe, 2019*
G. calviceps (Pocock, 1900)
G. childsi Kovarik, 2018*
G. dabakalo Kovarik & Mazuch, 2015*
G. gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013*
G. gubanensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. insolitus (Borelli, 1925)* (nomen dubium)
G. maidensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. marialuisae Rossi, 2015* (nomen dubium)
G. monicae Rossi, 2015* (nomen dubium)
G. puntlandus Kovarik & Mazuch, 2015*
|
Grosphus Simon,
1880 (14)
G. ambre Lourenco, Wilme & Waeber, 2018*
G. angulatus Lowe & Kovarik, 2022*
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. goudoti Lourenço & Goodman, 2006*
G. halleuxi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. hirtus Kraepelin, 1900
G. madagascariensis (Gervais, 1843)
G. mandena Lourenço, 2005*
G. mayottensis Lourenço & Goodman, 2009*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. rakotoariveloi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
G. tavaratra Lourenco, Soarimalala & Goodman, 2009*
G. voahangyae Lourenço & Wilme 2015*
|
Hemibuthus Pocock, 1900 (2)
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
|
Hemilychas Hirst,
1911 (1)
H. alexandrinus (Hirst, 1911)
|
Heteroctenus Pocock, 1893* (9)
H. abudi (Armas & Marcano Fondeur, 1987)*
H. aridicola (Teruel et Armas, 2012)*
H. bonettii (Armas 1999)*
H. garridoi (Armas, 1974)*
H. gibarae (Teruel, 2006)*
H. granulimanus (Teruel, 2006)*
H. junceus (Herbst, 1800)*
H. melloleitaoi (Teruel et Armas, 2006)*
H. princeps (Karsch, 1879)*
|
Himalayotityobuthus
Lourenço, 1997 (2)
H. alejandrae Lourenço, 2003*
H. martensi Lourenço,
1997
| |
Hottentotta
Birula, 1908 (60)
H. akbarii Yagmur, Moradi, Tabatabaei & Jafari, 2022*
H. alticola (Pocock, 1895)
H. alticola alticola (Pocock, 1895)
H. alticola nigrifrons (Pocock, 1900)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004* (nomen dubium)
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. flavidulus Teruel & Rein, 2010*
H. franzwerneri (Birula, 1914)
H. fuscitruncus (Caporiacco, 1936)
H. gambelaensis Kovarik, 2015*
H. gentili (Pallary, 1924)*
H. gibaensis Kovarik, 2015*
H. haudensis Kovarik & Lowe, 2021*
H. hoggarensis Lourenço & Leguin, 2014*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. juliae Kovarik, Yagmur & Fet, 2019*
H. keralaensis Aswhati, Sureshan & Lourenço, 2016*
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. krivokhatskyi Kovarik, Yagmur & Fet, 2019*
H. lacroixi Ythier & Dupre, 2021*
H. lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarik & Kayedi, 2010*
H. mazuchi Kovarik, 2013*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1875)
H. minax minax (L. Koch, 1975)
H. minax occidentalis (Vachon & Stockmann, 1968)
H. minusalta Vachon, 1959*
H. navidpouri Kovarik, Yagmur & Moradi, 2018*
H. nigrimontanus Kovarik & Lowe, 2021*
H. niloticus (Birula, 1928)*
H. novaki Kovarik, 2015*
H. pachyurus (Pocock, 1897)*
H. pellucidus Lowe, 2010*
H. penjabensis (Birula, 1897)*
H. polystictus (Pocock, 1896)
H. pooyani Moradi, Yagmur, Akbari & Jafari, 2022*
H. reddyi Lourenço, 2015*
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1880)
H. saxinatans Lowe, 2010*
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. sistanensis Kovarik, Yagmur & Moradi, 2018*
H. socotrensis (Pocock, 1889)
H. somalicus Kovarik, 2018*
H. songi (Lourenço, Qi & Zhu, 2005)*
H. sousai Turiel, 2014*
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trailini Kovarik, 2013*
H. trilineatus (Peters, 1861)
H. ugandaensis Kovarik, 2013*
H. vinchu Mirza, Ambekar & Kulkarni, 2019*
H. zagrosensis Kovarik, 1997
|
Iranobuthus
Kovarik, 1997 (1)
I. krali Kovarik,
1997
|
Ischnotelson Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (2)
I. guanambiensis (Lenarducci, Pinto-da-Rocha & Lucas, 2005)*
I. peruassu Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
|
Isometroides
Keyserling, 1885 (1)
I. vescus (Karsch,
1880)
|
Isometrus Ehrenberg,
1828 (14)
I. amboli Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. atherii Amir & Kamaluddin, 2008*
I. formosus Pocock, 1894
I. isadensis Tikader & Bastawade, 1983*
I. kovariki Sulakhe, Dandekar, Mukherjee, Pandey, Ketkar, Padhye & Bastawade, 2020*
I. liaqatii Amir & Kamaluddin, 2008*
I. longitelson Deshpande, Gowande, Bastawade & Sulakhe, 2022*
I. maculatus (DeGeer, 1778)
I. nakshatra Sulakhe, Deshpande, Gowande, Dandekar & Ketkar, 2022*
I. sankeriensis Tikander & Bastawade, 1983
I. tamhini Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. thurstoni Pocock, 1893
I. thwaitesi Pocock, 1897
I. wayanadensis Sulakhe, Deshpande, Gowande, Dandekar & Ketkar, 2022*
|
Jaguajir Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (3)
J. agamemnon (C. L. Koch, 1839)*
J. pintoi (Mello-Leitao, 1932)*
J. pintoi pintoi (Mello-Leitao, 1932)*
J. pintoi kourouensis (Lourenco, 2008)*
J. rochae (Borelli, 1910)*
|
Janalychas Kovarik, 2019* (9)
J. albimanus (Henderson, 1919)
J. farkasi (Kovarik, 1997)
J. granulatus Mirza, 2020*
J. heurtaultae (Kovarik, 1997)
J. keralaensis Mirza, 2020*
J. laevifrons (Pocock, 1897)
J. shoplandi (Oates, 1888)
J. srilankensis (Lorenco, 1997)
J. tricarinatus (Simon, 1884)
|
Karasbergia
Hewitt, 1913 (1)
K. methueni Hewitt, 1913 |
Kraepelinia
Vachon, 1974 (1)
K. palpator (Birula, 1903) |
Lanzatus Kovarik,
2001* (3)
L. huluul Kovarik & Lowe, 2021*
L. somalicus Kovarik, 2001*
L. somalilandus Kovarik & Lowe, 2016* |
Leiurus Ehrenberg, 1828 (20)
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. aegyptiacus Lourenço & El-Hennawy, 2021*
L. arabicus Lowe, Yagmur & Kovarik, 2014*
L. ater Lourenço, 2019*
L. brachycentrus (Ehrenberg, 1829)*
L. dekeyseri Lourenço, 2020*
L. gubanensis Kovarik & Lowe, 2020*
L. haenggii Lowe, Yagmur & Kovarik, 2014*
L. heberti Lowe, Yagmur & Kovarik, 2014*
L. hebraeus (Birula, 1908)*
L. hoggarensis Lourenço, Kourim & Sadine, 2018*
L. kuwaiti Lourenço, 2020*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. macroctenus Lowe, Yagmur & Kovarik, 2014*
L. maculatus Lourenço, 2022*
L. nigerianus Lourenço, 2021*
L. quinquestriatus (Ehrenberg, 1828)
L. saharicus Lourenço, 2020*
L. savanicola Lourenço, Qi & Cloudsley-Thompson, 2006*
L. somalicus Lourenço, & Rossi, 2016*
| |
Liobuthus Birula,
1898 (1)
L. kessleri Birula, 1898 |
Lissothus Vachon, 1948 (3)
L. bernardi Vachon, 1948
L. chaambi Lourenço & Sadine, 2014*
L. occidentalis Vachon, 1950 |
Lychas C.L. Koch,
1845 (33)
L. aberlenci Lourenço, 2013*
L. armasi Kovarik, 2013*
L. armillatus (Gervais, 1841)*
L. asper (Pocock, 1891)
L. biharensis Tikader & Bastawade, 1983
L. brehieri Lourenço, 2017*
L. buchari Kovarik, 1997
L. cernickai Kovarik, 2013*
L. chanthaburiensis Ythier & Lourenco, 2022*
L. flavimanus (Thorell,
1888)
L. gravelyi Henderson, 1913
L. hendersoni (Pocock, 1897)
L. hillyardi Kovarik, 1997
L. inexpectatus Lourenço, 2011*
L. jakli Kovarik, 2023*
L. kamshetensis Tikader & Bastawade, 1983
L. kharpadi Bastawade, 1986
L. krali Kovarik, 1995
L. lourencoi Kovarik, 1997
L. marmoreus (C.L. Koch, 1844)
L. mjobergi Kraepelin, 1916
L. mucronatus (Fabricius, 1798)
L. nigristernis (Pocock, 1899)
L. obsti Kraepelin, 1913
L. perfidus (Keyserling, 1885)
L. rackae Kovarik, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. variatus (Thorell, 1876)
L. variatus variatus (Thorell, 1876)
L. variatus canopensis Lourenco & Qi, 2007
|
|
Lychasioides
Vachon, 1974 (1)
L. amieti Vachon, 1974 |
Mauritanobuthus Qi & Lourenço, 2007* (1)
M. geniezi Qi &Lourenço, 2007*
|
Mesobuthus Vachon, 1950 (29)
M. afghanus (Pocock, 1889)*
M. barszczewskii (Birula, 1904)
M. birulai Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. bogdoensis (Birula, 1896)*
M. crucittii Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. eupeus (C.L. Koch, 1839)
M. fomichevi Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. farleyi Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. galinae Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. haarlovi Vachon, 1958*
M. iranus (Birula, 1917)*
M. kaftani Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. kirmanensis (Birula, 1900)*
M. macmahoni (Pocock, 1900)
M. marusiki Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. mesopotamicus Penther, 1912*
M. mirshamsii Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. navidpouri Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. persicus (Pocock, 1899)*
M. philippovitschi (Birula, 1905)*
M. phillipsi (Pocock, 1889)*
M. rahsenae Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. thersites (C. L. Koch, 1839)*
M. turcicus Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. vesiculatus (Pocock, 1899)
M. vignolii Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. yagmuri Kovarik & Fet, 2022*
M. zarudnyi Nouvruzov, Kovarik & Fet, 2022*
M. zonsteini Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
|
Mesotityus
Gonzalez-Sponga, 1981 (1)
M. vondangeli
Gonzalez-Sponga, 1981 |
Microananteris
Lourenço, 2003* (4)
M. abounami Lourenço & Chevalier, 2022*
M. inselberg Lourenço, 2021*
M. minor Lourenço, 2003*
M. serrulata Lourenço, 2021*
|
Microbuthus
Kraepelin, 1898 (7)
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. gardneri Lowe, 2010*
M. kristensenorum Lowe, 2010*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
M. satyrus Lowe, Kovarik, Stockmann & Stahlavsky, 2018*
|
Microcharmus Lourenço, 1995 (16)
M. andrei Lourenço, Waeber & Wilme, 2019*
M. antongil Lourenço, Waeber & Wilme, 2019*
M. bemaraha Lourenço, Goodman & Fisher, 2006*
M. cloudsleythompsoni Lourenço, 1995
M. confluenciatus Lourenço, Goodman & Fisher, 2006*
M. djangoa Lourenço, Waeber & Wilme, 2019*
M. duhemi Lourenço, Goodman & Fisher, 2006*
M. fisheri Lourenço, 1998
M. hauseri Lourenço, 1996
M. jussarae Lourenço, 1996*
M. maculatus Lourenço, Goodman & Fisher, 2006*
M. madagascariensis Lourenço, 1999*
M. pauliani (Lourenço, 2004)*
M. pauliani pauliani (Lourenço, 2004)
M. pauliani ambre Lourenco, Goodman & Fisher, 2006
M. pauliani namoroka Lourenco, Goodman & Fisher, 2006
M. sabineae Lourenço, 1996
M. variegatus Lourenço, Goodman & Fisher, 2006*
M. violaceous Lourenço, Goodman & Fisher, 2006*
|
Microtityus
Kjellesvig-Waering, 1966 (40)
M. angelaerrosae Gonzales-Sponga, 2001*
M. barahona Armas & Teruel, 2012*
M. biordi Gonzalez-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. borincanus Teruel, Rivera & Sanchez, 2014*
M. capayaensis Gonzalez-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae Gonzalez-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. eustatia Armas, 2018*
M. farleyi Teruel, 2000*
M. flavescens Teruel, 2001*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M. jaumei Armas, 1974
M. joseantonioi Gonzalez-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis Gonzalez-Sponga, 2001*
M. lourencoi Armas & Teruel, 2012*
M. minimus Kovarik & Teruel, 2014*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. prendinii Armas & Teruel, 2012*
M. pusillus Teruel & Kovarik, 2012*
M. reini Armas & Teruel, 2012*
M. rickyi Kjellesvig-Waering, 1966
M. santosi Teruel, Rivera & Sanchez, 2014*
M. sevciki Gonzalez-Sponga, 2001*
M. solegladi Armas & Teruel, 2012*
M. starri Lourenço & Huber, 1999*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. vieques Teruel, Rivera & Santos, 2015*
M. virginiae Armas, 1999*
M. vulcanicus Teruel, 2019*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus Gonzalez-Sponga, 2001*
|
Neobuthus Hirst,
1911 (16)
N. amoudensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. awashensis Kovarik & Lowe, 2012*
N. berberensis Hirst, 1911
N. cloudsleythompsoni Lourenço, 2001*
N. erigavoensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. eritreaensis Lowe & Kovarik, 2016*
N. factorio Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. ferrugineus (Kraepelin, 1898)*
N. gubanensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. haeckeli Kovarik, 2019*
N. kloppersi Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. kutcheri Lowe & Kovarik, 2016*
N. maidensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. montanus Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. solegladi Kovarik, 2019*
N. sudanensis Lourenço, 2005*
|
Neogrosphus Lourenço, 1995 (3)
N. andrafiabe Lourenço, Wilme & Waeber 2015*
N. blanci Lourenço, 1996
N. griveaudi (Vachon, 1969) |
Neoprotobuthus
Lourenço, 2000* (1)
N. intermedius Lourenço,
2000* |
Odontobuthus Vachon, 1950 (9)
O. baluchicus Barahoei, Prendini, Navidpour, Tahir, Aliabadian & Siahsarvie, 2021*
O. bidentatus Lourenço & Pezier, 2002*
O. brevidigitus Lowe, 2010*
O. chabaharensis Barahoei, Prendini, Navidpour, Tahir, Aliabadian & Siahsarvie, 2021*
O. doriae (Thorell, 1876)
O. kermanus Barahoei, Prendini, Navidpour, Tahir, Aliabadian & Siahsarvie, 2021*
O. odonturus (Pocock, 1897)
O. tavighiae Navidpour, Soleglad, Fet & Kovarik, 2013*
O. tirgari Mirshamsi, Azghadi, Navidpour, Aliabadian & Kovarik, 2013*
|
Odonturus Karsch,
1879 (1)
O.
dentatus Karsch, 1879 |
Olivierus Farzanpay, 1987 (21)
O. bolensis (Sun, Zhu & Lourenço, 2010)*
O. brutus (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. caucasicus (Nordmann, 1840)
O. elenae (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. extremus (Werner, 1936)
O. fuscus (Birula, 1897)*
O. gorelovi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. hainanensis (Birula, 1904)
O. intermedius (Birula, 1897)*
O. karshius (Sun & Sun, 2011)*
O. kaznakovi (Birula, 1904)*
O. kreuzbergi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. longichelus (Sun & Zhu, 2010)*
O. martensii (Karsch, 1879)
O. mikhailovi Fet, Kovarik, Gantenbein & Graham, 2021*
O. mischi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. nenilini (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. parthorum (Pocock, 1889)*
O. przewalskii (Birula, 1897)
O. tarabaevi Fet, Kovarik, Gantenbein & Graham, 2021*
O. voldemari Fet, Kovarik, Gantenbein & Graham, 2021*
|
Orthochiroides
Kovarik, 1998 (4)
O. insularis (Pocock, 1899)*
O. socotrensis Kovarik, 2004*
O. somalilandus Kovarik & Lowe, 2022*
O. vachoni Kovarik, 1998
|
Orthochirus Karsch, 1891 (59)
O. afar Kovarik & Lowe, 2016*
O. afghanus Kovarik, 2004*
O. arabicus Ythier & Lourenço, 2023*
O. arenicola Lourenço & Ythier, 2021* [nomen dubium]
O. atarensis Lourenço & Leguin, 2011*
O. bastawadei Zambre, Mirza, Sanap, Upadhye & Javed, 2011*
O. bicolor (Pocock, 1897)
O. birulai Kovarik, Fet & Yagmur, 2020*
O. carinatus Navidpour, Kovarik, Soleglad & Fet, 2019*
O. cloudsleythompsoni Lourenço & Leguin, 2011*
O. danielleae (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fomichevi Kovarik, Yagmur, Fet & Hussen, 2019*
O. formozovi Kovarik, Fet & Yagmur, 2020*
O. fuscipes (Pocock, 1900)
O. gantenbeini Kovarik, Yagmur, Fet & Hussen, 2019*
O. glabrifrons (Kraepelin, 1903)*
O. gromovi Kovarik, 2004*
O. grosseri Kovarik, Fet & Yagmur, 2020*
O. gruberi Kovarik & Fet, 2006*
O. hormozganensis Kovarik & Navidpour, 2020*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. kaspareki (Lourenço & Huber, 2000)*
O. katerinae Kovarik & Just, 2022*
O. kermanensis Kovarik & Navidpour, 2020*
O. kinzelbachi (Lourenço & Huber, 2000)*
O. kovariki Yagmur & Khalili, 2022*
O. krishnai Tikader & Bastawade, 1983
O. kryzhanovskyi Kovarik, Fet & Yagmur, 2020*
O. kucerai Kovarik & Navidpour, 2020*
O. masihipouri Kovarik & Navidpour, 2020*
O. maroccanus Lourenço & Leguin, 2011*
O. melanurus (Kessler, 1874)*
O. mesopotamicus Birula, 1918*
O. milloti Lourenço, 2021*
O. minor Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. monodi (Lourenço & Vachon, 1997)*
O. navidpouri Kovarik, Yagmur, Fet & Hussen, 2019*
O. nordmanni Kovarik, Fet & Yagmur, 2020*
O. olivaceus (Karsch, 1881)
O. pallidus (Pocock, 1897)
O. persa (Birula, 1900)*
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. scrobiculosus scrobiculosus (Grube, 1873)
O. scrobiculosus negebensis (Shulov & Amitai, 1960)
O. sejnai Kovarik, Fet & Yagmur, 2020*
O. semnanensis Kovarik & Navidpour, 2020*
O. soufiensis Lourenco & Sadine, 2021*
O. stockwelli (Lourenço & Vachon, 1995)*
O. tassili Lourenço & Leguin, 2011*
O. tibesti Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. varius Kovarik, 2004*
O. vignolii Kovarik & Navidpour, 2020*
O. zagrosensis Kovarik, 2004*
|
Pantobuthus
Lourenço & Duhem, 2009* (1)
P. complicatus Lourenço & Duhem, 2009*
|
Parabuthus Pocock,
1890 (38)
P. abyssinicus Pocock, 1901*
P. brevimanus (Thorell, 1876)
P. calvus Purcell, 1898
P. capensis (Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral, 1980
P. erigavoensis Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. eritreaensis Kovarik, 2003*
P. glabrimanus Prendini & Esposito, 2010*
P. gracilis Lamoral, 1979
P. granimanus Pocock, 1895
P. granulatus (Ehrenberg, 1831)
P. hamar Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. heterurus Pocock, 1899
P. hunteri Pocock, 1895
P. kabateki Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. kajibu Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. kalaharicus Lamoral, 1977
P. kraepelini Werner, 1902
P. kuanyamarum Monard, 1937
P. laevifrons (Simon, 1888)
P. liosoma (Ehrenberg, 1828)
P. maximus Werner, 1913
P. mazuchi Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. mossambicensis (Peters, 1861)
P. muelleri Prendini, 2000*
P. namibensis Lamoral, 1979
P. nanus Lamoral, 1979
P. pallidus Pocock, 1895
P. planicauda (Pocock, 1889)
P. raudus (Simon, 1888)
P. robustus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. schlechteri Purcell, 1899
P. setiventer Prendini & Esposito, 2010*
P. somalilandus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. stridulus Hewitt, 1913
P.
transvaalicus Purcell, 1899
P.
villosus (Peters, 1862)
|
Pectinibuthus Fet in Orlov & Vasilyev, 1984 (1)
P. birulai Fet in Orlov & Vasilyev, 1984 |
Physoctonus
Mello-Leitao, 1934* (3)
P. amazonicus Lourenço, 2017*
P. debilis (C. L. Koch, 1840)*
P. striatus Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
|
Picobuthus Lowe, 2010* (2)
P. dundoni Lowe, 2010*
P. wahibaensis Lowe, 2010*
|
Plesiobuthus
Pocock, 1900 (1)
P. paradoxus Pocock, 1900
|
Polisius Fet,
Capes & Sissom, 2001* (1)
P. persicus Fet,
Capes & Sissom, 2001* |
|
Pseudolissothus
Lourenço, 2001* (1)
P. pusillus Lourenço, 2001*
|
Pseudolychas
Kraepelin, 1911 (3)
P. ochraceus (Hirst,
1911)
P. pegleri (Purcell, 1901)
P.
transvaalicus Lawrence, 1961
|
|
Pseudouroplectes
Lourenço, 1995 (6)
P. betschi Lourenço, 1995
P. jacki Lourenço, 2021*
P. lalyae Lourenço & Ythier, 2010*
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999*
P. tsingy Lourenço, Wilme & Waeber, 2016*
|
Razianus Farzanpay, 1987 (4)
R. birulai Tahir, Navidpour & Prendini, 2014*
R. farzanpayi Tahir, Navidpour & Prendini, 2014*
R. xinjianganus Lourenco, Sun & Zhu, 2010*
R. zarudnyi (Birula, 1903)
|
Reddyanus Vachon, 1972* (34)
R. aareyensis (Mirza & Sanap, 2010)*
R. acanthurus (Pocock, 1899)*
R. assamensis (Oates, 1888)*
R. basilicus (Karsch, 1879)*
R. besucheti (Vachon, 1982)*
R. bilyi (Kovarik, 2003)*
R. brachycentrus (Pocock, 1899)*
R. ceylonensis Kovarik, Lowe, Ranawana, Hoferek, Jayarathne, Pliskova & Stahlavsky, 2016*
R. corbeti (Tikander & Batawade, 1983)*
R. deharvengi (Lourenço & Duhem, 2010)*
R. feti (Kovarik, 2013)*
R. furai Kovarik & Stahlavsky, 2019*
R. heimi (Vachon, 1976)*
R. hofereki Kovarik & Stahlavsky, 2019*
R. jayarathnei Kovarik, 2016*
R. jendeki (Kovarik, 2013)*
R. justi Kovarik, Lowe & Stahlavsky, 2020*
R. khammamensis (Kovarik, 2003)*
R. krasenskyi (Kovarik, 1998)*
R. kurkai (Kovarik, 1997)*
R. loebli (Vachon, 1982)*
R. majkusi Kovarik & Stahlavsky, 2019*
R. melanodactylus (L. Koch, 1867)*
R. navaiae (Kovarik, 1998)*
R. neradi (Kovarik, 2013)*
R. petrzelkai (Kovarik, 2003)*
R. problematicus (Kovarik, 2003)*
R. ranawanai Kovarik, 2016*
R. rigidulus (Pocock, 1897)*
R. rolciki Kovarik & Stahlavsky, 2019*
R. schwotti Kovarik & Stahlavsky, 2019*
R. tibetanus (Lourenço & Zhu, 2008)*
R. vittatus (Pocock, 1900)
R. zideki (Kovarik, 1994)*
|
|
Rhopalurus Thorell, 1876 (3)
R. caribensis Teruel & Roncallo, 2008*
R. laticauda Thorell, 1876
R. ochoai Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
|
|
|
|
Saharobuthus
Lourenço & Duhem, 2009* (1)
S. elegans Lourenço & Duhem, 2009*
|
Sassanidotus
Farzanpay, 1987 (2)
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900)
|
Somalibuthus
Kovarik, 1998 (2)
S. demisi Kovarik, 1998
S. sabae Kovarik & Njoroge, 2021*
|
Somalicharmus
Kovarik, 1998 (1)
S. whitmanae Kovarik, 1998 |
Spelaeolychas Kovarik, 2019* (1)
S. hosei (Pocock, 1891)
|
Teruelius Lowe & Kovarik, 2019* (22)
T. ankarafantsika (Lourenço, 2003)*
T. ankarana (Lourenço & Goodman, 2003)*
T. annulatus (Fage, 1929)*
T. bemaraha (Lourenco, Wilme & Waeber, 2018)*
T. bicolor (Lourenço, 2012)*
T. bistriatus (Kraepelin, 1900)*
T. eliseanneae (Lourenço & Wilme, 2016)*
T. feti (Lourenço, 1996)*
T. flavopiceus (Kraepelin, 1900)*
T. ganzhorni (Lourenço, Wilme & Waeber, 2016)*
T. grandidieri (Kraepelin, 1900)*
T. haeckeli Lowe & Kovarik, 2022*
T. intertidalis (Lourenço, 1999)*
T. limbatus (Pocock, 1889)*
T. magalieae (Lourenço, 2014)*
T. mahafaliensis (Lourenço, Goodman & Ramilijaona, 2004)*
T. makay (Lourenço & Wilme, 2015)*
T. mavo (Lourenço & Rossi 2020)*
T. olgae (Lourenço, 2004)*
T. rossii (Lourenço, 2013)*
T. sabineae (Lourenço & Wilme, 2016)*
T. waeberi (Lourenço & Wilme, 2016)*
|
Thaicharmus Kovarik, 1995 (4)
T. guptai Mirza, Sanap & Kunte, 2016*
T. indicus Kovarik, 2013*
T. lowei Kovarik, Soleglad & Fet, 2007*
T. mahunkai Kovarik, 1995
|
|
Tityobuthus Pocock, 1893 (21)
T. antsingy Lourenço & Goodman, 2004*
T. baroni (Pocock, 1890)
T. betschi Lourenço, Qi & Goodman, 2008*
T. chelbergorum Lourenço, Qi & Goodman, 2008*
T. darainensis Lourenço & Goodman, 2002*
T. dastychi Lourenço, 1997
T. griswoldi Lourenço, 2000*
T. guillaumeti Lourenço, 1995
T. ivohibe Lourenço & Goodman, 1999*
T. judsoni Lourenço, 1996
T. lokobe Lourenço, Waeber & Wilme, 2016*
T. manonae Lourenço, 2000*
T. mariejeanneae Lourenço, Waeber & Wilme, 2018*
T. mccarteri Lourenço, Qi & Goodman, 2008*
T. monodi Lourenço, 2000*
T. orangea Lourenço, Waeber & Wilme, 2020*
T. pallidus Lourenço, 2004*
T. parrilloi Lourenço, 1996
T. petrae Lourenço, 1996
T. pococki Lourenço, 1995
T. rakotondravonyi Lourenço & Goodman, 2003*
|
Tityopsis Armas,
1974 (6)
T. canizaresorum Teruel & Rodriguez-Cabrera, 2020*
T. inaequalis (Armas, 1974)
T. inexpectata (Moreno, 1940)
T. mulata Teruel & Rodriguez-Cabrera, 2020*
T. pumila Teruel & Rodriguez-Cabrera, 2020*
T. sheylae Teruel & Rodriguez-Cabrera, 2020*
|
Tityus C. L. Koch, 1836 (227)
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. abudi Armas, 1999*
T. acananensis Gonzalez-Sponga, 2009*
T. adisi Lourenço & Pezier, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi Gonzalez-Sponga, 2001*
T. alejandroi Teruel, Rivera & Santos, 2015*
T. altithronus Armas, 1999*
T. androcottoides (Karsch, 1879)
T. anduzei Gonzalez-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. anori Lourenço, Rossi & Wilme, 2019*
T. antioquensis Lourenço& Otero Patino, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai Gonzalez-Sponga, 1985
T. argentinus Borelli, 1899
T. asthenes Pocock, 1893
T. atriventer Pocock, 1897
T. bahiensis (Perty, 1833)
T. bahiensis bahiensis (Perty, 1833)
T. bahiensis eickstedtae Lourenço, 1982
T. barquisimetanus Gonzalez-Sponga, 1994
T. bastosi Lourenço, 1984
T. bellulus Armas, 1999*
T. birabeni Abalos, 1955
T. blanci Lourenço, 1994
T. blaseri Mello-Leitao, 1931
T. boconoensis Gonzalez-Sponga, 1981
T. bolivianus Kraepelin, 1895
T. brazilae Lourenço & Eickstedt, 1984
T. breweri Gonzalez-Sponga, 1997
T. cachipalensis Gonzalez-Sponga, 2002*
T. caesarbarrioi Gonzalez-Sponga, 2001*
T. canopensis Lourenço & Pezier, 2002*
T. carabobensis Gonzalez-Sponga, 1987
T. carinatoides Mello-Leitao, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carolineae Kovarik, Teruel, Cozijn & Seiter, 2013
T. carrilloi Ojanguren-Affilastro, 2021*
T. carvalhoi Mello-Leitao, 1945
T. cerroazul Lourenço, 1986
T. championi Pocock, 1898*
T. charalaensis Mello-Leitao, 1940 [nomen nudum]
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. choco Lourenço & Florez D, 2018*
T. cisandinus Lourenço & Ythier, 2017*
T. clathratus C. L. Koch, 1844
T. columbianus (Thorell, 1876)
T. confluens Borelli, 1899
T. confluens confluens Borelli, 1899
T. confluens bodoquena Lourenço, Cabral & Bruehmueller Ramos, 2004
T. costatus (Karsch, 1879)
T. crassicauda Lourenço & Ythier, 2013*
T. crassimanus (Thorell, 1876)
T. cuellari Lourenço, 1994
T. culebrensis Gonzalez-Sponga, 1994
T. curupi Ojanguren-Affilastro, Adilardi, Cajade, Ramirez, Ceccarelli & Mola, 2017*
T. cylindricus (Karsch, 1879)*
T. dasyurus Pocock, 1897
T. dasyurus dasyurus Pocock, 1897
T. dasyurus fulvipes Mello-Leitao, 1945
T. dedoslargos Francke & Stockwell, 1987
T. demangei Lourenço, 1981
T. dillerorum Kovarik, Teruel, Lowe & Friedrich, 2015*
T. dinizi Lourenço, 1997
T. discrepans (Karsch, 1879)
T. dorae Gonzalez-Sponga, 2001*
T. duacaensis Gonzalez-Sponga, 2007*
T. dulceae Gonzalez-Sponga, 2006*
T. dupouyi Gonzalez-Sponga, 1987
T. ecuadorensis
Kraepelin, 1896
T. elii Armas & Marcano
Fondeur, 1992*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi Gonzalez-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leitao, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessoa, 1935
T. festae Borelli, 1899
T. filodendron Gonzalez-Sponga, 1981
T. florezi Lourenço, 2000*
T. footei Chamberlin, 1916
T. forcipula (Gervais, 1843)
T. fuhrmanni Kraepelin, 1914
T. funestus Hirst, 1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. generaltheophiloi Lourenço, 2017*
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. grahami Lourenço, 2012*
T. grottoedensis Botero-Trujillo & Florez D, 2014*
T. guane Moreno-Gonzalez, Gonzalez O & Florez D, 2019*
T. guaricoensis Gonzalez-Sponga, 2004*
T. haetianus Teruel & Santos, 2018*
T. horacioi Lourenço& Leguin, 2011*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus Mello-Leitao, 1934
T. insignis (Pocock, 1889)
T. intermedius Borelli, 1899
T. isabelceciliae Gonzalez-Sponga, D'Suze & Sevcik, 2001*
T. ivani Gonzalez-Sponga, 2008*
T. ivicnancor Gonzalez-Sponga,
1997
T. jaimei Miranda, Bermudez, Florez D & de Armas, 2020*
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai Gonzalez-Sponga, 2007*
T. karaja Lourenço, 2016*
T. kindli Kovarik & Teruel, 2014*
T. kukututee Ythier, Chevalier & Gangadin, 2020*
T. kuryi Lourenço, 1997
T. lancinii Gonzalez-Sponga, 1972
T. lokiae Lourenço, Adis & Araujo, 2005*
T. longidigitus Gonzalez-Sponga, 2008*
T. lourencoi Florez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis Gonzalez-Sponga, 2007*
T. mana Lourenço, 2012*
T. manakai Gonzalez-Sponga, 2004*
T. maniapurensis Gonzalez-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. marechali Lourenço, 2013*
T. martinpaechi Lourenço, 2001*
T. matthieseni Pinto-da-Rocha & Lourenço, 2000
T. mattogrossensis Borelli,
1901
T. maturinensis Gonzalez-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis Gonzalez-Sponga, 1974
T. mongei Lourenço, 1996
T. moralensis Moreno-Gonzalez, Pinto da Rocha & Cabra-Garcia, 2022*
T. mucusunamensis Gonzalez-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leitao, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leitao, 1940
T. neoespartanus Gonzalez-Sponga, 1996
T. nororientalis Gonzalez-Sponga, 1996
T. obispoi Gonzalez-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus Gonzalez-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis Gonzalez-Sponga, 2007*
T. panguana Kovarik, Teruel, Lowe & Friedrich, 2015*
T. paraguayensis Kraepelin, 1895
T.
parvulus Kraepelin, 1914
T. paulistorum Lourenço & da Silva, 2006*
T.
perijanensis Gonzalez-Sponga, 1994
T.
pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri
Gonzalez-Sponga, 1981
T. pococki Hirst,
1907
T. portoplatensis Armas & Marcano
Fondeur, 1992
T. potameis Lourenço & Leao Giupponi, 2004*
T. prancei Lourenço,
2000*
T. proseni Abalos, 1954
T.
pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis Gonzalez-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus Armas, 1982*
T. ramirezi Esquivel de Verde, 1969 [nomen dubium]
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis Gonzalez-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi Gonzalez-Sponga, 1996
T. romeroi Gonzalez-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rosenbergi Pocock, 1898*
T. rufofuscus Pocock, 1897
T. rupestre Lourenço 2019*
T. rugosus Schenkel, 1932
T. rusmelyae Gonzalez-Sponga, D'Suze & Sevcik, 2001*
T. sabineae Lourenço, 1994
T. sanarensis Gonzalez-Sponga, 1997
T. sanfernandoi Gonzalez-Sponga, 2008*
T. sarisarinamensis Gonzalez-Sponga, 2002*
T. sastrei Lourenço & Florez, 1990
T. schrammi Teruel & Santos, 2018*
T. serrulatus Lutz & Mello, 1922
T. shiriana Gonzalez-Sponga, 1991
T. silvestris Pocock, 1897
T. simonsi Pocock, 1900
T. smithii Pocock, 1893
T. soratensis Kraepelin, 1911
T. spelaeus Moreno-Gonzalez, Pinto-da-Rocha & Gallao, 2021*
T. stigmurus (Thorell, 1876)
T. strandi Werner, 1939
T. surmeridensis Gonzalez-Sponga, 2002*
T. surorientalis Gonzalez-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi Gonzalez-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. timendus Pocock, 1898
T. trinitatis Pocock, 1897
T. trivittatus Kraepelin, 1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leitao, 1931
T. unus Pinto-da-Rocha & Lourenço, 2000
T. uquirensis Gonzalez-Sponga, 2001*
T. urachichensis Gonzalez-Sponga, 2007*
T. urbinai
Scorza, 1952
T. uruguayensis Borelli,
1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis Gonzalez-Sponga, 1981
T. ventuarensis Gonzalez-Sponga, 2009*
T. wachteli Kovarik, Teruel, Lowe & Friedrich, 2015*
T. walli Gonzalez-Sponga & Wall Gonzalez, 2007*
T. yerenai Gonzalez-Sponga, 2009*
T. ythieri Lourenço, 2007*
T. zulianus Gonzalez-Sponga, 1981
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Troglorhopalurus Lourenço, Baptista & Giupponi,
2004* (2)
T. lacrau (Lourenço & Pinto-da-Rocha, 1997)*
T. translucidus Lourenço, Baptista & Giupponi, 2004*
| Troglotityobuthus Lourenço,
2000* (1)
T. gracilis (Fage, 1946)
|
Trypanothacus Lowe, Kovarik, Stockmann & Stahlavsky, 2019* (3)
T. azraqensis Al-Saraireh, Afifeh, Aloufi, Amr & Lourenco, 2021*
T. barnesi Lowe, Kovarik, Stockmann & Stahlavsky, 2019*
T. buettikeri (Hendrixson, 2006)*
| |
Uroplectes Peters,
1861 (34)
U. ansiedippenaarae Prendini, 2015*
U. carinatus (Pocock, 1890)
U. carinatus carinatus (Pocock, 1890)
U. carinatus australis Hewitt, 1918
U. chubbi Hirst, 1911
U. emiliae (Werner, 1916)
U. fischeri (Karsch, 1879)
U. flavoviridis Peters, 1861
U. formosus Pocock, 1890
U. formosus formosus Pocock, 1890
U. formosus basuticus Hewitt, 1927
U. formosus maculipes Hewitt, 1918
U. formosus spenceri Pocock, 1896
U. gracilior Hewitt, 1913
U. insignis Pocock, 1890
U. katangensis Prendini, 2015*
U. lineatus (C. L. Koch, 1844)
U. longimanus Werner, 1936
U. machadoi Lourenço, 2000*
U. malawicus Prendini, 2015*
U. marlothi Purcell, 1901
U. ngangelarum Monard, 1930
U. occidentalis Simon, 1876
U. occidentalis occidentalis Simon, 1876
U. occidentalis monardi Vachon, 1950
U. olivaceus Pocock, 1896
U. otjimbinguensis (Karsch, 1879)
U. otjimbinguensis otjimbinguensis (Karsch, 1879)
U. otjimbinguensis massacarum Monard, 1937
U. pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner, 1913
U. pilosus (Thorell, 1876)
U. planimanus (Karsch, 1879)
U. planimanus planimanus (Karsch, 1879)
U. planimanus kuanyamarus Monard, 1937
U. schlechteri Purcell, 1901
U. schubotzi Kraepelin, 1929
U. silvestrii Borelli, 1913
U. teretipes Lawrence, 1966
U. triangulifer (Thorell, 1876)
U. tumidimanus Lamoral, 1979
U. variegatus (C. L. Koch, 1844)
U. vittatus (Thorell, 1876)
U. xanthogrammus Pocock, 1897
U. zambezicus Prendini, 2015*
| |
Vachoniolus Levy, Amitai & Shulov, 1973 (4)
V. batinahensis Lowe, 2010*
V. gallagheri Lowe, 2010*
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
|
Vachonus Tikader & Bastawade, 1983 (4)
V. asiyaae Amir & Kamaluddin, 2009* [nomen dubium]
V. inexpectatus Lourenço, 2015*
V. iqbali Amir & Kamaluddin, 2009* [nomen dubium]
V. rajasthanicus Tikader & Bastawade, 1983
|
Xenobuthus Lowe, 2018* (3)
X. anthracinus (Pocock, 1895)*
X. arabicus (Lourenço & Qi, 2006)*
X. xanthus Lowe, 2018*
|
Zabius Thorell, 1893 (3)
Z. birabeni Mello-Laeitao, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*
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