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CHACTIDAE
Pocock, 1893 |
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This family consists of 11 genera and 168 species [Updated 01.03.10].
Most species have been described in recent times. There
has been a lot of discussion about some of the genera and
species in this family, and several genera have been
transferred to other families.
In a recent revision by Soleglad & Fet, 36 species previously listed in Broteochactas
were transfered to a new genus, Neochactas. The genera Cayooca, Guyanochactas and Taurepania have been
synonymized with Broteochactas. Three species from Broteochactas have been transfered to Hadrurochactas. The same paper transfered the European genus Belisarius to Chactidae (this genus was previously listed in the discontinued family Troglotayosicidae), but I've now reinstated Troglotayosicidae and moved the genus back again (see Troglotayosicidae for more info).
In addition, the genus Uroctonus has been transfered from the family Vaejovidae.
The members of this family are
known from North America, Central- and South America.
Soleglad & Sissom, 2001 have
transfered the genus Chactopsis to the family Euscorpiidae.
There and back again: Soleglad & Fet (2005) have restored the synonymization of
Auyantepuia with Broteochactas after Lourenço & Araujo (2004)
reinstated the genus Auyantepuia (which before this was synonymized with Broteochactas (and later
some species with Neochactas)). Of the 8 species of Auyantepuia, one is transfered to Broteochactas and
the others are transfered back to Neochactas (se the species list below for details).
Lourenço & Qi, 2007 have published a new species in Auyantepuia, and stating that they do not accept the work of
Soleglad & Fet (2005) (Citing Prendini & Wheeler, 2005). Auyantepuia is reinstated in the species list to accomodate the
new species until a new revision of the family is published.
Scorpions in this family are rarely kept in captivity.
Chactid scorpions are not medically important.
Brotheas granulatus photo by
Dr. Wilson Lourenço (C) and The Journal of Venomous Animals and
Toxins/CEVAP (C)
SPECIES
FILES:
This list of genera and
species is based on Fet
et al.(2000). Subspecies is not included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000)
See also Soleglad & Fet (2003) for more information. Fulltext freely available HERE (issue 11).
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Anuroctonus Pocock, 1893
A. pococki Soleglad & Fet, 2004*
A. phaiodactylus (Wood, 1863)
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[Auyantepuia (Gonzalez-Sponga, 1978)*]
A. amapaensis Lourenço & Qi, 2007*
As described in the introduction, the genus Auyantepuia has been synonymized with other genera.
Lourenço & Qi (2007) have chosen not to accept this synonymization, and described the new species in Auyantepuia. The taxonomy of
The Scorpion Files follows Soleglad & Fet (2005), but it is impossible for me to know where to put the new species. I have chosen to reinstate Auyantepuia in The
Scorpion Files for this species until a new revision on the family Chactidae is published. Auyantepuia is not counted in
the number of genera for the family, but the species is included. |
Broteochactas
Pocock, 1893
B. cocuyensis Gonzalez-Sponga, 2004*
B. danielleae Lourenço, 2007*
B. gollmeri (Karsch, 1879)
B. gonzalezspongai (Lourenço, 1983)
B. goujei Vellard, 1932
B. manisapanensis (González-Sponga, 1992)
B. mascarenhasi (Lourenço, 1983)
B. nitidus Pocock, 1893
B. parimensis González-Sponga, 2004*
B. porosus (Pocock, 1900)
B. scorzai Dagert, 1957
B. trezzii(Vignoli & Kovarik, 2003)*
B. venezuelensis (González-Sponga, 1996)
B. verneti (González-Sponga, 1992)
B. vestigialis González-Sponga, 1978
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Brotheas
C.L. Koch, 1837
B. amazonicus Lourenço, 1988
B. bolivianus (Lourenço, 2008)*
B. camposi González-Sponga, 1972
B. cataniapensis González-Sponga, 1997
B. cristinae Lourenço, 2007*
B. cunucunumensis González-Sponga,
1984
B. dasilvai González-Sponga,
1978
B. gervaisii Pocock, 1893
B.
granimanus Pocock, 1898
B. granulatus
Simon, 1877
B. henriquesi Lourenço & Machado, 2004*
B. humboldti González-Sponga,
1980
B. jourdani Lourenço, 1997
B.
libinallyi González-Sponga, 1978
B.
lichyi González-Sponga, 1980
B.
mawarinumensis González-Sponga, 1991
B.
mingueti González-Sponga, 1973
B.
munozi González-Sponga, 1997
B. ocamoi González-Sponga, 2004*
B. overali
Lourenço, 1988
B. paraensis Simon,
1880
B. perezramirezi González-Sponga,
1996
B. rionegroensis González-Sponga,
1996
B. sanabriai González-Sponga,
1997
B. silvestris Lourenço, 1988
B. wareipai González-Sponga, 2004*
B.
wilmeri González-Sponga, 1980
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Chactas
Gervais, 1844
C.
aequinoctialis (Karsch, 1879)
C. alarconi González-Sponga, 2003*
C. barbacoensis González-Sponga, 1987
C. barravierai Lourenço, 1997
C. braziliensis Lourenço, Aguiar & Franklin, 2005*
C. brevicaudatus (Karsch, 1879)
C. brownelli Lourenço, 1997
C. campoelisasensis González-Sponga, 2006*
C. camposi Mello-Leităo, 1939
C. chabasquensis González-Sponga & Wall-González, 2007*
C. choroniensis González-Sponga, 1978
C. chrysopus Pocock, 1893
C. exsul (Werner, 1939)
C. ferruginosus González-Sponga, 1984
C. gansi González-Sponga, 1974
C. gestroi Kraepelin, 1912
C. granulosus González-Sponga, 2007*
C. guinandcortesi González-Sponga, 2003*
C. hatilloensis González-Sponga, 2007*
C. hauseri Lourenço, 1997
C. interpuncta González-Sponga, 1987
C. iutensis González-Sponga, 2008*
C. karschii Pocock, 1893
C. keyserlingii Pocock, 1893
C. koepckei Lourenço & Dastych, 2001*
C. laevipes (Karsch, 1879)
C. latuffi González-Sponga, 1976
C. lepturus Thorell, 1867
C. major Kraepelin, 1914*
C. mahnerti Lourenço, 1995
C. maimirensis González-Sponga & Wall-González, 2007*
C. marinae González-Sponga, 1987
C. mauriesi Lourenço & Florez, 1990
C. ozendai Lourenço, 1999*
C. oxfordi González-Sponga, 1978
C. platillonensis González-Sponga & Wall-González, 2007*
C. raymondhansi Francke & Boos, 1986
C. reticulatus Kraepelin, 1912
C. rogelioi González-Sponga, 1972
C. rubrolineatus Simon, 1880
C. scabrimanus Kraepelin, 1912
C. setosus Kraepelin, 1912
C. simonii Pocock, 1893
C. tumaquensis González-Sponga & Wall-González, 2007*
C. vanbenedenii (Gervais, 1843)
C. venegasi González-Sponga, 2008*
C. viloriai Rojas-Runjaic, 2004*
C. turguaensis González-Sponga, 2007*
C. yupai González-Sponga, 1994 |
Hadrurochactas
Pocock, 1893
H. araripe (Lourenço, 2010*)
H. brejo (Lourenço, 1988)
H. machadoi González-Sponga, 1993*
H. mapuera (Lourenço, 1988)
H. odoardoi González-Sponga, 1985
B. polisi (Monod & Lourenço, 2001)*
H. schaumii (Karsch,
1880)
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Neochactas Soleglad & Fet 2003*
N. bariensis (González-Sponga, 1991)
N. bilbaoi (González-Sponga, 1978)
N. bruzuali (González-Sponga, 1980)
N. caroniensis (González-Sponga, 1996)
N. colombiensis (González-Sponga, 1976)
N. delicatus (Karsch, 1879)
N. efreni (González-Sponga, 1978)
N. eliasilvai (González-Sponga, 1980)
N. fei (Pinto-da-Rocha, Gasnier, Brescovit & Apolinario, 2002)*
N. fravalae (Lourenço, 1983)*
N. gaillardi (Lourenço,1983)
N. garciai (González-Sponga, 1978)
N. granosus (Pocock, 1900)
N. guaiquinimensis (González-Sponga, 1997)
N. jaspei (Gonzalez-Sponga, 1993)
N. josemanueli (González-Sponga, 1992)
N. kelleri (Lourenço, 1997)
N. kjellesvigi (González-Sponga, 1974)
N. laui (Kjellesvig-Waering, 1966)
N. leoneli (González-Sponga, 1978)
N. macrochelae (González-Sponga, 2004)*
N. mottai (Lourenço & Araujo, 2004)*
N. neblinensis (González-Sponga, 1991)
N. orinocensis (Scorza, 1954)
N. panarei (González-Sponga, 1980)
N. paoensis (González-Sponga, 1996)
N. parvulus (Pocock, 1897)
N. racenisi (González-Sponga, 1975)
N. riopinensis (González-Sponga, 1992)
N. ruizpittoli (González-Sponga, 1993)
N. sanmartini (González-Sponga, 1974)
N. santanai (González-Sponga, 1978)
N. sarisarinamensis (González-Sponga, 1985)
N. simarawochensis (González-Sponga, 1980)
N. sissomi (Lourenço, 1983)
N. skuki (Lourenço & Pinto-da-Rocha, 2000)*
N. verai (González-Sponga, 1993)
N. yekuanae (González-Sponga, 1984)
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Nullibrotheas
Williams, 1974
N. allenii
(Wood, 1863) |
Teuthraustes
Simon, 1878
T. adrianae
González-Sponga, 1975
T. akananensis
González-Sponga, 1984
T. amazonicus (Simon,
1880)
T. atramentarius Simon, 1878
T.
carmelinae Scorza, 1954
T. dubius
(Borelli, 1899)
T. festae (Borelli,
1899)
T. gervaisii (Pocock, 1893)
T.
glaber Kraepelin, 1912
T. guerdouxi
Lourenço, 1995
T. lisei Lourenço,
1994
T. lojanus (Pocock, 1900)
T.
maturaca González-Sponga, 1991
T.
oculatus Pocock, 1900
T. ohausi
Kraepelin, 1912
T. reticulatus
González-Sponga, 1991
T. rosenbergi (Pocock,
1898)
T. simonsi (Pocock, 1900)
T.
whymperi (Pocock, 1893)
T. wittii
(Kraepelin, 1896)
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Uroctonus Thorell, 1876
U. franckei Williams, 1986
U. grahami Gertsch & Soleglad, 1972
U. mordax Thorell, 1876
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Vachoniochactas
Gonzalez-Sponga, 1978
V. amazonicus González-Sponga, 1991
V. ashleeae Lourenço, 1994
V. humboldti Florez, Botero-Trujillo & Acosta, 2008*
V. lasallei (González-Sponga, 1978)
V. roraima Lourenço & Duhem, 2009
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