Akravidae
Bothriuridae
Buthidae
Caraboctonidae
Chactidae
Chaerilidae
Euscorpiidae
Hemiscorpiidae
Heteroscorpionidae
Hormuridae
Iuridae
Pseudo-
chactidae
Scorpionidae
Superstitioniidae
Troglotayosicidae
Typhlochactidae
Vaejovidae
About the family
systematics

BUTHIDAE
C.L. Koch, 1837

With 90 genera (one extinct and not listed below) and 1016 species [Updated 07.03.14], this family is the largest of the scorpion families. This family is widespread around the world (not found in Antarctica and New Zealand), and are found in tropical, subtropical and partly in temperate habitats.

Volschenk, Mattoni & Prendini (2008) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus and Neoprotobuthus) and 15 species are all transfered to Buthidae. Information about the old family Microcharmidae.

Kovarik (2007) has published a revision of the genus Hottentotta. Note that the medical important Mesobuthus tamulus and two other Asian Mesobuthus have been transfered to Hottentotta. Kovarik's revision can be downloaded HERE (browse down to issue 58).

Paraorthochirus has been synonymized with Orthochirus by Navidpour et al., 2008).

The members of this family are small and medium sized scorpions with a triangular sternum (some genera have a more pentagonal sternum). In many genera (e.g., Androctonus, Apistobuthus, Parabuthus, and other Old World genera) the cauda is usually strong and powerful and the pedipalps are often very slender whereas other genera have elongate appendages, especially the males (e.g., Centruroides, Lychas, Isometrus and related genera). Many species are yellow or brown (or variations of these colors), but black forms are also present. Some species have quite intricate and attractive patterns (e.g., Lychas and Isometrus) and colours (e.g., Centruroides and Uroplectes). Sizes range from about 20 mm (like Microtityus and Microbuthus) to over 120 mm (as in Androctonus, some Centruroides, Apistobuthus and others).

An indentification key to the genus Compsobuthus was published by Kovarik (2012) [Free full text].

Several species in this family are highly toxic, but fewer than 20 can be lethal to man. The most notorious species are found in the genus Androctonus, Centruroides, Hottentotta, Leiurus, Parabuthus and Tityus. See Scorpions of medical importance page for further information.

Unlike most other scorpion families, the Buthidae are of both scientific and public interest due to their medical importance in many parts of the world. Several species have also made their way into the hobby. Many of these are safe to be kept by experienced collectors, but the most dangerous species should be avoided.

Androctonus crassicauda habitat picture from Kuwait are located HERE.

Photo of Parabuthus leiosoma (left) by Jan Ove Rein (C)
Photo of Hottentotta sp. (right) by Jan Ove Rein (C)

SPECIES FILES:
This list of genera and species is based on Fet et al.(2000). Subfamilies and subspecies is not included in the list. I try to update the list as additions and changes are published. * denotes changes after Fet et al. (2000). I will be grateful for information about new development in the taxonomy of this family. For information about synonyms and bibliographies, see Fet et al. (2000)

Afghanobuthus Lourenço, 2005*
A. naumanni Lourenço, 2005*
Afroisometrus Kovarík, 1997
A. minshullae (Fitzpatrick, 1994)
Akentrobuthus Lamoral, 1976
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976
Alayotityus Armas, 1973
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A. juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas, 1973

Ananteris Thorell, 1891
A. arcadioi Botero-Trujillo, 2008*
A. ashmolei Lourenço, 1981
A. asuncionensis González-Sponga, 2006*
A. balzanii Thorell, 1891
A. barinensis González-Sponga, 2006*
A. bernabei Giupponi, Vasconcelos & Lourenço, 2009*
A. bianchinii Lourenço, Aguiar-Neto & Limeira-de-Oliveira, 2009*
A. bonito Lourenço, 2012*
A. cachimboensis Lourenço, Motta & da Silva, 2006*
A. camacan Lourenço, Giupponi & Leguin 2013*
A. capayaensis González-Sponga, 2006*
A. caracensis González-Sponga, 2006*
A. catuaroi González-Sponga, 2006*
A. caucaguitensis González-Sponga, 2006*
A. chagasi Giupponi, Vasconcelos & Lourenço, 2009*
A. charlescorfieldi Lourenço, 2001*
A. chirimakei González-Sponga, 2006*
A. claviformis González-Sponga, 2006*
A. coinaui Lourenço, 1982
A. columbiana Lourenço, 1991
A. cryptozoicus Lourenço, 2005*
A. cumbensis González-Sponga, 2006*
A. curariensis González-Sponga, 2006*
A. cussinii Borelli, 1910
A. dekeyseri Lourenço, 1982
A. deniseae Lourenço, 1997
A. desiderio Lourenço, Giupponi & Leguin 2013*
A. diegorojasi Rojas-Runjaic, 2005*
A. dorae Botero-Trujillo, 2008*
A. ehrlichi Lourenço, 1994
A. elguapoi González-Sponga, 2006*
A. elisabethae Lourenço, 2003*
A. evellynae Lourenço, 2004*
A. faguasi Botero-Trujillo, 2009*
A. festae Borelli, 1899
A. franckei Lourenço, 1982
A. gorgonae Lourenço & Flórez, 1989
A. guiripaensis González-Sponga, 2006*
A. guyanensis Lourenço & Monod, 1999*
A. infuscata Lourenço, Giupponi & Leguin 2013*
A. inoae González-Sponga, 2006*
A. intermedia Lourenço, 2012*
A. kuryi Giupponi, Vasconcelos & Lourenço, 2009*
A. leilae Lourenço, 1999*
A. luciae Lourenço, 1984
A. madeirensis Lourenço & Duhem, 2010*
A. maniapurensis González-Sponga, 2006*
A. maranhensis Lourenço, 1987
A. mariaelenae Lourenço, 1999*
A. mariaterezae Lourenço, 1987
A. mauryi Lourenço, 1982
A. meridana González-Sponga, 2006*
A. michaelae Lourenço, 2013*
A. myriamae Botero-Trujillo, 2007*
A. nairae Lourenço, 2004*
A. norae González-Sponga, 2006*
A. ochoai Botero-Trujillo & Florez D, 2011*
A. palmari Botero-Trujillo & Noriega, 2011*
A. paoensis González-Sponga, 2006*
A. paracotoensis González-Sponga, 2006*
A. plataensis González-Sponga, 2006*
A. platnicki Lourenço, 1993
A. principalis González-Sponga, 2006*
A. pydanieli Lourenço, 1982
A. riocaurensis González-Sponga, 2006*
A. riochicoi González-Sponga, 2006*
A. riomachensis Rojas-Runjaic, Portillo-Quintero & Borges, 2008*
A. roraima Lourenço & Duhem, 2010*
A. sabineae Lourenço, 2001*
A. sanchezi González-Sponga, 2006*
A. sepulvedai González-Sponga, 2006*
A. singularis González-Sponga, 2006*
A. solimariae Botero-Trujillo & Florez D, 2011*
A. surinamensis Lourenço, 2012*
A. terueli Kovarík, 2006*
A. tolimana Teruel & Garcia, 2007*
A. turumbanensis Gonzáles-Sponga, 1990
A. venezuelensis Gonzáles-Sponga, 1972
A. volschenki Botero-Trujillo, 2009*
A. zuliana González-Sponga, 2006*

Ananteroides Borelli, 1911*
A. feae Borelli, 1911*
A. inexpectatus Lourenço, 2013*
 
Androctonus Ehrenberg, 1828
A. afghanus Lourenço & Qi, 2006*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi (Audouin,1826)
A. australis (Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. bicolor Ehrenberg, 1828
A. cholistanus Kovarik & Ahmed, 2013*
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Yhtier & Leguin, 2009*
A. pallidus Lourenço, Duhem & Cloudsley-Thompson, 2012*
A. robustus Kovarik & Ahmed, 2013*
A. sergenti Vachon, 1948
A. tenuissimus Teruel, Kovarik & Turiel, 2013*
A. togolensis Lourenço, 2008*
Anomalobuthus Kraepelin, 1900
A. rickmersi Kraepelin, 1900
Apistobuthus Finnegan, 1932
A. pterygocercus Finnegan, 1932
A. susanae Lourenço, 1998
Australobuthus Locket, 1990 A. xerolimniorum Locket, 1990
Babycurus Karsch, 1886
B. ansorgei Hirst, 1911
B. buettneri Karsch, 1886
B. centrurimorphus Karsch, 1886*
B. exquisitus Lowe, 2000*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. ornatus Werner, 1936
B. pictus Pocock, 1896*
B. prudenti Lourenço, 2013*
B. solegladi Lourenço, 2005*
B. somalicus Hirst, 1907
B. subpunctatus Borelli, 1925
B. ugartei Kovarik, 2000*
B. wituensis Kraepelin, 1913
B. zambonellii Borelli, 1902
Baloorthochirus Kovarik, 1996
B. becvari Kovarík, 1996
Birulatus Vachon, 1974
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
Buthacus Birula, 1908
B. agarwali Zambre & Lourenço, 2010*
B. arenicola (Simon, 1885)
B. armasi Lourenço, 2013*
B. birulai Lourenço, 2006*
B. buettikeri Hendrixson, 2006*
B. clevai Lourenço, 2001*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (Nomen dubium)
B. golovatchi Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. huberi Lourenço, 2001*
B. leptochelys (Ehrenberg, 1829)
B. macrocentrus (Ehrenberg, 1828)*
B. mahraouii Lourenço, 2004*
B. maliensis Lourenço & Qi, 2007*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. occidentalis Lourenço, 2000*
B. pakistanensis Lourenço & Qi, 2006*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. willamsi Lourenço & Leguin, 2009*
B. ziegleri Lourenço, 2000*
Butheoloides Hirst, 1925
B. annieae Lourenço, 1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. granulatus Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. hirsti Lourenço, 1996
B. littoralis Lourenço, Touloun & Boumezzough, 2011*
B. maroccanus Hirst, 1925
B. milloti Vachon, 1948
B. monodi Vachon, 1950
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. savanicola Lourenço, 2013*
B. schwendingeri Lourenço, 2002*
B. slimanii Lourenço, 2010*
B. wilsoni Lourenço, 1995
Butheolus Simon, 1882
B. andersoni (Pocock, 1895)*
B. anthracinus (Pocock, 1895)
B. arabicus Lourenço & Qi, 2006*
B. gallagheri Vachon, 1980
B. pallidus Lourenço & Duhem, 2012*
B. thalassinus Simon, 1882
B. villosus Hendrixson, 2006*
Buthiscus Birula, 1905
B. bicalcaratus Birula, 1905
Buthoscorpio Werner, 1936
B. indicus Lourenço, 2012*
B. politus (Pocock, 1899)
B. rayalensis Javed, Rao, Mirza, Sanap & Tampal, 2010*
B. sarasinorum (Karsch, 1891)
NB! B. jinnahii Amir, Kamaluddin & Jabbar, 2005* and B. rahmatii Amir, Kamaluddin & Jabbar, 2005* was described in the non-valid genus Stenochirus by the authors. I have placed them in Buthoscorpio, which is the valid name for Stenochirus, but formally this has to be done in a scientific journal before it is valid. Javed et al., 2010 have studied the descriptions of these two species and concluded that they do not belong to the genus Buthoscorpio. This decsision was also supported by Lourenco, 2012. The two taxa are now Buthidae incertae sedis and are not included in the total number species in Buthidae.
Buthus Leach, 1815
B. adrianae Rossi, 2013
B. albengai Lourenço, 2003*
B. amri Lourenço, Yagmur & Duhem 2010*
B. atlantis Pocock, 1889
B. awashensis Kovarik, 2011*
B. barcaeus Birula, 1909*
B. barbouri Werner, 1932
B. berberensis Pocock, 1900*
B. bonito Lourenço & Geniez, 2005*
B. boumalenii Touloun & Boumezzough, 2011*
B. brignolii Lourenço, 2003*
B. chambiensis Kovarik, 2006*
B. confluens Lourenço, Touloun & Boumezzough, 2012*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. egyptiensis Lourenço, 2012*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. elongatus Rossi, 2012*
B. hassanini Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. ibericus Lourenço & Vachon, 2004*
B. insolitus Borelli, 1925
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. israelis (Shulov & Amitai, 1959)*
B. jianxinae Lourenço, 2005*
B. kunti Yagmur, Koc & Lourenço, 2011*
B. lienhardi Lourenço, 2003*
B. lourencoi Rossi, Tropea & Yagmur, 2013*
B. malhommei Vachon, 1949*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. orientalis Lourenço & Simon, 2012*
B. paris (C. L. Koch, 1839)*
B. prudenti Lourenço & Leguin, 2012*
B. pusillus Lourenço, 2013*
B. rochati Lourenço, 2003*
B. tassili Lourenço, 2002*
B. trinacrius Lourenço & Rossi, 2013*+
+Probably extinct.
B. tunetanus (Herbst, 1800)*
B. yemenensis Lourenço, 2008*
Centruroides Marx, 1890
C. alayoni Armas, 1999*
C. anchorellus Armas, 1976
C. arctimanus (Armas, 1976)
C. baergi Hoffmann, 1932*
C. balsasensis Ponce & Francke, 2004*
C. bani Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis (Pocock, 1898)
C. bertholdii (Thorell, 1876)
C. bicolor (Pocock, 1898)
C. caral Armas & Trujillo. 2013
C. chamela Ponce-Saavedra & Francke, 2011*
C. chamulaensis Hoffmann, 1932
C. chiapanensis Hoffmann, 1932
C. edwardsii (Gervais, 1843)*
C. elegans (Thorell, 1876)
C. exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul (Meise, 1933)
C. fallassisimus Armas & Trujillo, 2010*
C. farri Armas, 1976
C. flavopictus (Pocock, 1898)
C. franckei Santibanez-Lopez & Contreras-Felix, 2013*
C. fulvipes (Pocock, 1898)
C. galano Teruel, 2001*
C. gracilis (Latreille, 1804)
C. granosus (Thorell, 1876)*
C. griseus (C. L. Koch, 1844)
C. guanensis Franganillo, 1930
C. hentzi (Banks, 1900)
C. hirsuticauda Teruel, 2011*
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. mahnerti Lourenço, 1983
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mariaorum Santiago-Blay, 2009*
C. mascota Ponce-Saavedra & Francke, 2011*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus (Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. platnicki Armas, 1981
C. pococki Sissom & Francke, 1983
C. polito Teruel, 2007*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. rodolfoi Santibanez-Lopez & Contreras-Felix, 2013*
C. sanandres Armas, Sarmiento & Florez 2012*
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. simplex (Thorell, 1876)*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. suffusus (Pocock, 1902)
C. tecomanus Hoffmann, 1932*
C. testaceus (DeGeer, 1778)
C. thorelli (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. villegasi Baldazo-Monsivaiz, Ponce-Saavedra & Flores-Moreno, 2013*
C. vittatus (Say, 1821)
Charmus Karsch, 1879
C. brignolii Lourenço, 2000*
C. indicus Hirst, 1915
C. laneus Karsch, 1879
C. minor Lourenço, 2002*
C. sinhagadensis Tikader & Bastawade, 1983
Cicileiurus Teruel, 2007*
C. monticola Teruel, 2007*
Cicileus Vachon, 1948
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
Compsobuthus Vachon, 1949
C. abyssinicus (Birula, 1903)
C. acutecarinatus (Simon, 1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov, 1973
C. atrostriatus (Pocock, 1897)*
C. berlandi Vachon, 1950
C. birulai Lourenço, Leguin & Duhem, 2010*
C. boucheti Lourenço, Duhem & Cloudsley-Thompson, 2012*
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. garyi Lourenço & Vachon, 2001*
C. humaae Amir, Kamaluddin & Kahn, 2005* (Nomen dubium)
C. jakesi Kovarik, 2003*
C. jordanensis Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. klaptoczi (Birula, 1909)
C. krali Kovarik, 2012*
C. levyi Kovarik, 2012*
C. longipalpis Levy, Amitai & Shulov, 1973
C. maindroni (Kraepelin, 1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. plutenkoi Kovarik, 2003*
C. polisi Lowe, 2001*
C. rugosulus (Pocock, 1900)
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. sindicus Kovarik & Ahmed, 2011*
C. somalilandus Kovarik, 2012*
C. tassili Lourenço, 2010*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. vachoni Sisson, 1994
C. werneri (Birula, 1908)
C. williamsi Lourenço, 1999*
Congobuthus Lourenço, 1999*
C. fagei Lourenço, 1999*
Darchenia Vachon, 1977
D. bernadettae Vachon, 1977
Egyptobuthus Lourenço, 1999*
E. vaissadei Lourenço, 1999*
Femtobuthus Lowe, 2010*
F. shutuae Lowe, 2010*
Gint Kovarik, Lowe, Pliskova & Stahlavsky, 2013*
G. calviceps (Pocock, 1900)
G. gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013*
Grosphus Simon, 1880
G. ankarafantsika Lourenço, 2003*
G. ankarana Lourenço & Goodman, 2003*
G. annulatus Fage, 1929
G. bicolor Lourenço, 2012*
G. bistriatus Kraepelin, 1900
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. feti Lourenço, 1996
G. flavopiceus Kraepelin, 1900
G. garciai Lourenço, 2001*
G. goudoti Lourenço & Goodman, 2006*
G. grandidieri Kraepelin, 1900
G. hirtus Kraepelin, 1900
G. intertidalis Lourenço, 1999*
G. limbatus (Pocock, 1889)
G. madagascariensis (Gervais, 1843)
G. mandena Lourenço, 2005*
G. mahafaliensis Lourenço, Goodman & Ramilijaona, 2004*
G. mayottensis Lourenço & Goodman, 2009*
G. olgae Lourenço, 2004*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. rossii Lourenço, 2013*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
Hemibuthus Pocock, 1900
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
Hemilychas Hirst, 1911
H. alexandrinus (Hirst, 1911)
Himalayotityobuthus Lourenço, 1997
H. alejandrae Lourenço, 2003*
H. martensi Lourenço, 1997
Hottentotta Birula, 1908
H. alticola (Pocock, 1895)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004* (Nomen dubium)
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. flavidulus Teruel & Rein, 2010*
H. franzwerneri (Birula, 1914)
H. fuscitruncus (Caporiacco, 1936)
H. gentili (Pallary, 1924)*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarik & Kayedi, 2010*
H. mazuchi Kovarik, 2013*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1975)
H. minusalta Vachon, 1959*
H. niloticus (Birula, 1928)*
H. pachyurus (Pocock, 1897)*
H. pellucidus Lowe, 2010*
H. penjabensis Kovarik, 2007*
H. polystictus (Pocock, 1896)
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1828)
H. saxinatans Lowe, 2010*
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. socotrensis (Pocock, 1889)
H. songi (Lourenço, Qi & Zhu, 2005)*
H. sousai Turiel, 2014*
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (Nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trailini Kovarik, 2013*
H. trilineatus (Peters, 1861)
H. ugandaensis Kovarik, 2013*
H. zagrosensis Kovarík, 1997
Iranobuthus Kovarík, 1997
I. krali Kovarík, 1997
Isometroides Keyserling, 1885
I. vescus (Karsch, 1880)
 
Isometrus Ehrenberg, 1828
I. acanthurus Pocock, 1899
I. assamensis Oates, 1888
I. atherii Amir & Kamaluddin, 2008*
I. basilicus Karsch, 1879
I. besucheti Vachon, 1982
I. bilyi Kovarík, 2003*
I. brachycentrus Pocock, 1899
I. corbeti Tikander & Batawade, 1983
I. deharvengi Lourenço & Duhem, 2010*
I. feti Kovarík, 2013*
I. formosus Pocock, 1894
I. garyi Lourenço & Huber, 2002*
I. heimi Vachon, 1976
I. isadensis Tikander & Bastawade, 1983
I. jendeki Kovarík, 2013*
I. khammamensis Kovarík, 2003*
I. krasenskyi Kovarík, 1998
I. kurkai Kovarík, 1997
I. liaqatii Amir & Kamaluddin, 2008*
I. loebli (Vachon, 1982)*
I. maculatus (DeGeer, 1778)
I. melanodactylus (L. Koch, 1867)
I. navaiae Kovarík, 1998
I. neradi Kovarík, 2013*
I. petrzelkai Kovarík, 2003*
I. problematicus Kovarík, 2003*
I. rigidulus Pocock, 1897
I. thurstoni Pocock, 1893
I. thwaitesi Pocock, 1897
I. tibetanus Lourenço & Zhu, 2008*
I. vittatus Pocock, 1900
I. zideki Kovarík, 1994
Karasbergia Hewitt, 1913
K. methueni Hewitt, 1913
Kraepelinia Vachon, 1974
K. palpator (Birula, 1903)
Lanzatus Kovarik, 2001*
L. somalicus Kovarik, 2001*
Leiurus Ehrenberg, 1828
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. nasheri Kovarik, 2007*
L. quinquestriatus (Ehrenberg, 1828)
L. savanicola Lourenço, Qi & Cloudsley-Thompson, 2006*
Liobuthus Birula, 1898
L. kessleri Birula, 1898
Lissothus Vachon, 1948
L. bernardi Vachon, 1948
L. occidentalis Vachon, 1950
Lychas C.L. Koch, 1845
L. aareyensis Mirza & Sanap, 2010*
L. aberlenci Lourenço, 2013*
L. albimanus Henderson, 1919
L. armasi Kovarík, 2013*
L. armillatus (Gervais, 1841)*
L. asper (Pocock, 1891)
L. biharensis Tikader & Bastawade, 1983
L. braueri (Kraepelin, 1986)
L. buchardi Kovarík, 1997
L. burdoi (Simon, 1882)
L. cernickai Kovarík, 2013*
L. farkasi Kovarik, 1997
L. flavimanus (Thorell, 1888)
L. gravelyi Henderson, 1913
L. hendersoni (Pocock, 1897)
L. heurtaultae Kovarík, 1997
L. hillyardi Kovarík, 1997
L. hosei (Pocock, 1891)
L. inexpectatus Lourenço, 2011*
L. kaimana Lourenço, 2011* (Nomen dubium)
L. kamshetensis Tikader & Bastawade, 1983
L. kharpadi Bastawade, 1986
L. krali Kovarík, 1995
L. laevifrons Pocock, 1897
L. Lourençoi Kovarík, 1997
L. marmoreus (C.L. Koch, 1844)
L. mjobergi Kraepelin, 1916
L. mucronatus (Fabricius. 1798)
L. nigristernis (Pocock, 1899)
L. obsti Kraepelin, 1913
L. perfidus (Keyserling, 1885)
L. rackae Kovarík, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. shoplandi (Oates, 1888)
L. srilankensis Lorenco, 1997
L. tricarinatus (Simon, 1884)
L. variatus (Thorell, 1876)
Lychasioides Vachon, 1974
L. amieti Vachon, 1974
Mauritanobuthus Qi & Lourenço, 2007*
M. geniezi Qi &Lourenço, 2007*
Mesobuthus Vachon, 1950
M. agnetis (Werner, 1936)
M. bolensis Sun, Zhu & Lourenço, 2010*
M. caucasicus (Nordmann, 1840)
M. cyprius Gantenbein & Kropf, 2000*
M. eupeus (C.L. Koch, 1839)
M. extremus (Werner, 1936)
M. gibbosus (Brulli, 1832)
M. karshius Sun & Sun, 2011*
M. longichelus Sun & Zhu, 2010*
M. macmahoni (Pocock, 1900)
M. martensii (Karsch, 1879)
M. nigrocinctus (Ehrenberg, 1828)*
M. phillipsi (Pocock, 1889)*
M. vesiculatus (Pocock, 1899)
Mesotityus Gonzalez-Sponga, 1981
M. vondangeli González-Sponga, 1981
Microananteris Lourenço, 2003*
M. minor Lourenço, 2003*
Microbuthus Kraepelin, 1898
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. gardneri Lowe, 2010*
M. kristensenorum Lowe, 2010*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
Microcharmus Lourenço, 1995
M. bemaraha Lourenço, Goodman & Fisher, 2006*
M. cloudsleythompsoni Lourenço, 1995
M. confluenciatus Lourenço, Goodman & Fisher, 2006*
M. duhemi Lourenço, Goodman & Fisher, 2006*
M. fisheri Lourenço, 1998
M. hauseri Lourenço, 1996
M. jussarae Lourenço, 1996*
M. maculatus Lourenço, Goodman & Fisher, 2006*
M. madagascariensis Lourenço, 1999*
M. pauliani (Lourenço, 2004)*
M. sabineae Lourenço, 1996
M. variegatus Lourenço, Goodman & Fisher, 2006*
M. violaceous Lourenço, Goodman & Fisher, 2006*
Microtityus Kjellesvig-Waering, 1966
M. angelaerrosae Gonzáles-Sponga, 2001*
M. barahona Armas & Teruel, 2012*
M. biordi González-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. borincanus Teruel, Rivera & Sanchez, 2014*
M. capayaensis González-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae González-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. farleyi Teruel, 2000*
M. flavescens Teruel, 2001*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M. jaumei Armas, 1974
M. joseantonioi González-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis González-Sponga, 2001*
M. lourencoi Armas & Teruel, 2012*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. prendini Armas & Teruel, 2012*
M. pusillus Teruel & Kovarik, 2012*
M. reini Armas & Teruel, 2012*
M. rickyi Kjellesvig-Waering, 1966
M. santosi Teruel, Rivera & Sanchez, 2014*
M. sevciki González-Sponga, 2001*
M. solegladi Armas & Teruel, 2012*
M. starri Lourenço & Huber, 1999*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. virginiae Armas, 1999*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus González-Sponga, 2001*
Neobuthus Hirst, 1911
N. awashensis Kovarik & Lowe, 2012*
N. cloudsleythompsoni Lourenço, 2001*
N. berberensis (Hirst, 1911)
N. ferrugineus (Kraepelin, 1898)*
N. sudanensis Lourenço, 2005*
Neogrosphus Lourenço, 1995
N. blanci Lourenço, 1996
N. griveaudi (Vachon, 1969)
Neoprotobuthus Lourenço, 2000*
N. intermedius Lourenço, 2000*
Odontobuthus Vachon, 1950
O. bidentatus (Lourenço & Pezier, 2002)*
O. brevidigitus Lowe, 2010*
O. doriae (Thorell, 1876)
O. odonturus (Pocock, 1897)
O. tavighiae Navidpour, Soleglad, Fet & Kovarik, 2013*
O. tirgari Mirshamsi, Azghadi, Navidpour, Aliabadian & Kovarik, 2013*
Odonturus Karsch, 1879
O. dentatus Karsch, 1879
Orthochiroides Kovarík, 1998
O. insularis (Pocock, 1889)*
O. socotrensis Kovarík, 2004*
O. vachoni Kovarík, 1998
Orthochirus Karsch, 1891
O. afghanus Kovarik, 2004*
O. aristidis (Simon, 1882)
O. atarensis Lourenço & Leguin, 2011*
O. bastawadei Zambre, Mirza, Sanap, Upadhye & Javed, 2011*
O. bicolor (Pocock, 1897)
O. blandini (Lourenço & Vachon, 1997)*
O. cloudsleythompsoni Lourenço & Leguin, 2011*
O. danielleae (Lourenço & Vachon, 1997)*
O. erardi (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fuscipes (Pocock, 1900)
O. glabrifrons(Kraepelin, 1903)*
O. gromovi Kovarik, 2004*
O. gruberi Kovarik & Fet, 2006*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. kaspareki (Lourenço & Huber, 2000)*
O. kinzelbachi (Lourenço & Huber, 2000)*
O. krishnai Tikader & Bastawade, 1983
O. maroccanus Lourenço & Leguin, 2011*
O. minor Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. monodi (Lourenço & Vachon, 1997)*
O. pallidus (Pocock, 1897)
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. stockwelli (Lourenço & Vachon, 1995)*
O. tassili Lourenço & Leguin, 2011*
O. tibesti Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. varius Kovarik, 2004*
O. zagrosensis Kovarik, 2004*
Pantobuthus Lourenço & Duhem, 2009*
P. complicatus Lourenço & Duhem, 2009*
Parabuthus Pocock, 1890
P. brevimanus (Thorell, 1876)
P. calvus Purcell, 1898
P. capensis (Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral, 1980
P. eritreaensis Kovarik, 2003*
P. glabrimanus Prendini & Esposito, 2010*
P. gracilis Lamoral, 1979
P. granimanus Pocock, 1895
P. granulatus (Ehrenberg, 1831)
P. heterurus Pocock, 1899
P. hunteri Pocock, 1895
P. kalaharicus Lamoral, 1977
P. kraepelini Werner, 1902
P. kuanyamarum Monrad, 1937
P. laevifrons (Simon, 1888)
P. liosoma (Ehrenberg, 1828)
P. maximus Werner, 1913
P. mossambicensis (Peters, 1861)
P. muelleri Prendini, 2000*
P. namibensis Lamoral, 1979
P. nanus Lamoral, 1979
P. pallidus Pocock, 1895
P. planicauda (Pocock, 1889)
P. raudus (Simon, 1888)
P. schlechteri Purcell, 1899
P. setiventer Prendini & Esposito, 2010*
P. stridulus Hewitt, 1913
P. transvaalicus Purcell, 1899
P. villosus (Peters, 1862)
Pectinibuthus Fet, 1984
P. birulai Fet, 1984
Physoctonus Mello-Leitao, 1934*
P. debilis (C. L. Koch, 1840)*
Picobuthus Lowe, 2010*
P. dundoni Lowe, 2010*
P. wahibaensis Lowe, 2010*
Plesiobuthus Pocock, 1900
P. paradoxus Pocock, 1900
Polisius Fet, Capes & Sissom, 2001*
P. persicus Fet, Capes & Sissom, 2001*
Psammobuthus Birula, 1911
P. zarudnyi Birula, 1911
Pseudolissothus Lourenço, 2001*
P. pussilus Lourenço, 2001*
Pseudolychas Kraepelin, 1911
P. ochraceus (Hirst, 1911)
P. pegleri (Purcell, 1901)
P. transvaalicus Lawrence, 1961
Pseudouroplectes Lourenço, 1995
P. betschi Lourenço, 1995
P. lalyae Lourenço & Ythier, 2010*
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999*
Razianus Farzanpay, 1987

R. xinjianganus Lourenco, Sun & Zhu, 2010*
R. zarudnyi (Birula, 1903)
Rhopalurus Thorell, 1876
R. abudi Armas & Marcano Fondeur, 1987
R. acromelas Lutz & Mello, 1922
R. agamemmon (C. L. Koch, 1839)
R. amazonicus Lourenço, 1986
R. aridicola Teruel & Armas, 2012*
R. bonettii Armas 1999*
R. caribensis Teruel & Roncallo, 2008*
R. crassicauda Caporiacco, 1947*
R. garridoi Armas, 1974
R. guanambiensis Lenarducci, Pinto-da-Rocha & Lucas, 2005*
R. gibarae Teruel, 2006*
R. granulimanus Teruel, 2006*
R. junceus (Herbst, 1800)
R. lacrau Lourenço & Pinto-da-Rocha, 1997
R. laticauda Thorell, 1876
R. melloleitaoi Teruel & Armas, 2006*
R. pintoi Mello-Leităo, 1932*
R. princeps (Karsch, 1879)
R. rochai Borelli, 1910
R. virkkii Santiago-Blay, 2009*
Riftobuthus Lourenço, Duhem & Cloudsley-Thompson 2010*
R. inexpectatus Lourenço, Duhem & Cloudsley-Thompson 2010*
Sabinebuthus Lourenço, 2001*
S. elegans Lourenço, 2001*
Saharobuthus Lourenço & Duhem, 2009*
S. elegans Lourenço & Duhem, 2009*
Sassandiothus Farzanpay, 1987
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900)
Somalibuthus Kovarík, 1998
S. demisi Kovarík, 1998
Somalicharmus Kovarík, 1998
S. withmanae Kovarík, 1998
Thaicharmus Kovarík, 1995
T. indicus Kovarík, 1995
T. lowei Kovarík, Soleglad & Fet, 2007*
T. mahunkai Kovarík, 1995
Tityobuthus Pocock, 1893
T. antsingy Lourenço & Goodman, 2004*
T. baroni (Pocock, 1890)
T. betschi Lourenço, Qi & Goodman, 2008*
T. chelbergorum Lourenço, Qi & Goodman, 2008*
T. darainensis Lourenço & Goodman, 2002*
T. dastychi Lourenço, 1997
T. griswoldi Lourenço, 2000*
T. guillaumeti Lourenço, 1995
T. ivohibe Lourenço & Goodman, 1999*
T. judsoni Lourenço, 1996
T. manonae Lourenço, 2000*
T. mccarteri Lourenço, Qi & Goodman, 2008*
T. monodi Lourenço, 2000*
T. pallidus Lourenço, 2004*
T. parrilloi Lourenço, 1996
T. petrae Lourenço, 1996
T. pococki Lourenço, 1995
T. rakotondravonyi Lourenço, 2003*
Tityopsis Armas, 1974
T. aliciae Armas & Frias, 1998*
T. inaequalis (Armas, 1974)
T. inexpectata (Moreno, 1940)
Tityus C.L. Koch, 1836
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. abudi Armas, 1999*
T. acananensis González-Sponga, 2009*
T. adisi Lourenço, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi González-Sponga, 2001*
T. altithronus Armas, 1999*
T. anasilviae Armas & Abud Antun, 2004*
T. androcottoides (Karsch, 1879)
T. anduzei González-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. antioquensis Lourenço& Otero Patińo, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai González-Sponga, 1985
T. argentinus Borelli, 1899
T. asthenes Pocock, 1893
T. atriventer Pocock, 1897
T. bahiensis (Perty, 1833)
T. bahoruco Teruel & Armas, 2006*
T. barquisimetanus González-Sponga, 1994
T. bastosi Lourenço, 1984
T. bellulus Armas, 1999*
T. betschi Lourenço, 1992
T. birabeni Abalos, 1955
T. blanci Lourenço, 1994
T. blaseri Mello-Leităo, 1931
T. boconoensis González-Sponga, 1981
T. bolivanus Kraepelin, 1895
T. brazilae Lourenço & Eickstedt, 1984
T. breweri González-Sponga, 1997
T. cachipalensis González-Sponga, 2002*
T. caesarbarrioi González-Sponga, 2001*
T. canopensis Lourenço & Pezier, 2002*
T. carabobensis González-Sponga, 1987
T. carinatoides Mello-Leităo, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carolineae Kovarik, Teruel, Cozijn & Seiter, 2013
T. carvalhoi Mello-Leităo, 1945
T. cerroazul Lourenço, 1986
T. championi Pocock, 1898*
T. charalaensis Mello-Leităo, 1940
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. clathratus C. L. Koch, 1844
T. columbianus (Thorell, 1876)
T. confluens Borelli, 1899
T. costatus (Karsch, 1879)

T. crassicauda Lourenço, 2013*
T. crassimanus (Thorell, 1876)
T. cuellari Lourenço, 1994
T. culebrensis González-Sponga, 1994
T. cylindricus (Karsch, 1879)*
T. dasyurus Pocock, 1897
T. dedoslargos Francke & Stockwell, 1987
T. demangei Lourenço, 1981
T. dinizi Lourenço, 1997
T. discrepans (Karsch, 1879)
T. dorae González-Sponga, 2001*
T. duacaensis González-Sponga, 2007*
T. dulceae González-Sponga, 2006*
T. dupouyi González-Sponga, 1987
T. ebanoverde Armas, 1999*
T. ecuadorensis Kraepelin, 1896
T. elii (Armas & Marcano Fondeur, 1992)*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi González-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leităo, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessôa, 1935
T. festae Borelli, 1899
T. filodendron González-Sponga, 1981
T. florezi Lourenço, 2000*
T. footei Chamberlin, 1916
T. forcipula (Gervais, 1843)
T. fuhrmanni Kraepelin, 1914
T. funestus Hirst, 1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. grahami Lourenço, 2012*
T. guariconesis Gonzalez-Sponga, 2004*
T. horacioi Lourenço& Leguin, 2011*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus Mello-Leităo, 1934
T. insignis (Pocock, 1889)
T. intermedius Borelli, 1899
T. isabelceciliae González-Sponga, D'Suze & Sevcik, 2001*
T. ivani González-Sponga, 2008*
T. ivicnancor González-Sponga, 1997
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai González-Sponga, 2007*
T. kuryi Lourenço, 1997
T. lancinii González-Sponga, 1972
T. lokiae Lourenço, 2005*
T. longidigitus González-Sponga, 2008*
T. Lourençoi Flórez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis González-Sponga, 2007*
T. mana Lourenço, 2012*
T. manakai González-Sponga, 2004*
T. maniapurensis González-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. marechali Lourenço, 2013*
T. martinpaechi Lourenço, 2001*
T. matthieseni Lourenço & Pinto-da-Rocha, 2000
T. mattogrossensis Borelli, 1901
T. maturinensis González-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis González-Sponga, 1974
T. mongei Lourenço, 1996
T. mucusunamensis González-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leităo, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leităo, 1940
T. neoespartanus González-Sponga, 1996
T. nororientalis González-Sponga, 1996
T. obispoi González-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus González-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis González-Sponga, 2007*
T. paraguayensis Kraepelin, 1895
T. parvulus Kraepelin, 1914
T. paulistorum Lourenço & Qi, 2006*
T. perijanensis González-Sponga, 1994
T. pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri González-Sponga, 1981
T. pococki Hirst, 1907
T. portoplatensis Armas & Marcano Fondeur, 1992
T. potameis Lourenço & Leăo Giupponi, 2004*
T. prancei Lourenço, 2000*
T. proseni Abalos, 1954
T. pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis González-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus (Armas, 1982)*
T. ramirezi Esquivel de Verde, 1969 [nomen dubium]
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis González-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi González-Sponga, 1996
T. romeroi González-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rufofuscus Pocock, 1897
T. rugosus Schenkel, 1932
T. rusmelyae González-Sponga, D'Suze & Sevcik, 2001*
T. sabinae Lourenço, 1994
T. sanarensis González-Sponga, 1997
T. sanfernandoi González-Sponga, 2008*
T. sarisarinamensis González-Sponga, 2002*
T. sastrei Lourenço & Flórez, 1990
T. septentrionalis Armas & Abud Antun, 2004*
T. serrulatus Lutz & Mello, 1922
T. shiriana González-Sponga, 1991
T. silvestris Pocock, 1897
T. simonsi Pocock, 1900
T. smithii Pocock, 1893
T. soratensis Kraepelin, 1912
T. stigmurus (Thorell, 1876)
T. strandi Werner, 1939
T. surimeridensis González-Sponga, 2002*
T. surorientalis González-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi González-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. thelyacanthus Mello-Leităo, 1933
T. timendus Pocock, 1898
T. trinitatis Pocock, 1897
T. trivittatus Kraepelin, 1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leităo, 1931
T. unus Lourenço & Pinto-da-Rocha, 2000
T. uquirensis González-Sponga, 2001*
T. urachichensis González-Sponga, 2007*
T. urbinai Scorza, 1952
T. uruguayensis Borelli, 1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis González-Sponga, 1981
T. ventuarensis González-Sponga, 2009*
T. walli González-Sponga & Wall Gonzalez, 2007*
T. wayuu Rojas-Runjaic & Armas, 2007*
T. yerenai González-Sponga, 2009*
T. ythieri Lourenço, 2007*
T. zulianus González-Sponga, 1981
Troglorhopalurus Lourenço, Baptista & Giupponi, 2004*
T. translucidus Lourenço, Baptista & Giupponi, 2004*
Troglotityobuthus Lourenço, 2000*
T. gracilis (Fage, 1946)
Uroplectes Peters, 1861
U. fischeri (Karsch, 1879)
U. andreae Pocock, 1889
U. carinatus (Pocock, 1890)
U. chubbi Hirst, 1911
U. fischeri (Karsch, 1879)
U. flavoviridis Peters, 1861
U. formosus Pocock, 1890
U. gracilior Hewitt, 1914
U. insignis Pocock, 1890
U. lineatus (C. L. Koch, 1844)
U. longimanus Werner, 1936
U. marlothi Purcell, 1901
U. ngangelarum Monard, 1930
U. occidentalis Simon, 1876
U. olivaceus Pocock, 1896
U. otjimbinguensis (Karsch, 1879)
U. pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner, 1913
U. pilosus (Thorell, 1876)
U. planimanus (Karsch, 1879)
U. schlechteri Purcell, 1901
U. schubotzi Kraepelin, 1929
U. silvestrii Borelli, 1913
U. teretipes Lawrence, 1966
U. triangulifer (Thorell, 1876)
U. tumidimanus Lamoral, 1979
U. variegatus (C. L. Koch, 1844)
U. vittatus (Thorell, 1876)
U. xanthogrammus Pocock, 1897
Uroplectoides Lourenço, 1998
U. abyssinicus Lourenço, 1998
U. emiliae (Werner, 1916)
Vachoniolus Levy, Amitai & Shulov, 1973
V. batinahensis Lowe, 2010*
V. gallagheri Lowe, 2010*
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
Vachonus Tikader & Bastawade, 1983
V. asiyaae Amir & Kamaluddin, 2009*
V. iqbali Amir & Kamaluddin, 2009*
V. rajasthanicus Tikader & Bastawade, 1983
Zabius Thorell, 1893
Z. birabeni Mello-Laeităo, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*

Jan Ove Rein (C) 2013