| With 85 genera (one extinct) and 895 species [Updated 27.01.10],
this family is the largest of the scorpion families. This
family is widespread around the world (not found in Antarctica
and New Zealand), and are found in tropical, subtropical and
partly in temperate habitats.
Volschenk, Mattoni & Prendini (2008) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus and Neoprotobuthus) and 15 species are all transfered to Buthidae. Information about the old family Microcharmidae.
Kovarik (2007) has published a revision of the genus Hottentotta. Note that
the medical important Mesobuthus tamulus and two other Asian Mesobuthus have been
transfered to Hottentotta. Kovarik's revision can be
downloaded HERE (browse down to issue 58).
Paraorthochirus has been synonymized with Orthochirus by Navidpour et al., 2008).
The members of this family are small and medium sized
scorpions with a triangular sternum (some genera have a more
pentagonal sternum). In many genera (e.g., Androctonus,
Apistobuthus, Parabuthus, and other Old World
genera) the cauda is usually strong and powerful and the
pedipalps are often very slender whereas other genera have
elongate appendages, especially the males (e.g.,
Centruroides, Lychas, Isometrus and
related genera). Many species are yellow or brown (or
variations of these colors), but black forms are also present.
Some species have quite intricate and attractive patterns
(e.g., Lychas and Isometrus) and colours (e.g.,
Centruroides and Uroplectes). Sizes range from
about 20 mm (like Microtityus and Microbuthus)
to over 120 mm (as in Androctonus, some
Centruroides, Apistobuthus and others).
Several species in this family are highly toxic, but fewer
than 20 can be lethal to man. The most notorious species are
found in the genus Androctonus, Centruroides, Hottentotta,
Leiurus, Parabuthus and Tityus. See Scorpions
of medical importance page for further information.
Unlike most other scorpion families, the Buthidae are of
both scientific and public interest due to their medical
importance in many parts of the world. Several species have
also made their way into the hobby. Many of these are safe to
be kept by experienced collectors, but the most dangerous
species should be avoided.
Androctonus crassicauda habitat picture from Kuwait are located HERE.
Photo of Parabuthus leiosoma (left)
by Jan Ove Rein (C)
Photo of Hottentotta sp.
(right) by Jan Ove Rein (C)
SPECIES
FILES:
This list of genera and
species is based on Fet
et al.(2000). Subfamilies and subspecies is not included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000) |
Afghanobuthus
Lourenço, 2005*
A. naumanni Lourenço, 2005*
|
|
Afroisometrus
Kovarík, 1997
A. minshullae (Fitzpatrick,
1994)
|
Akentrobuthus
Lamoral, 1976
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976 |
Alayotityus Armas,
1973
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A.
juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus
Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas,
1973
|
Ananteris Thorell,
1891
A. arcadioi Botero-Trujillo, 2008*
A. ashmolei Lourenço, 1981
A.
asuncionensis González-Sponga, 2006*
A.
balzanii Thorell, 1891
A.
barinensis González-Sponga, 2006*
A. bernabei Giupponi, Vasconcelos & Lourenço, 2009*
A. bianchinii Lourenço, Aguiar-Neto & Limeira-de-Oliveira, 2009*
A. cachimboensis Lourenço, Motta & da Silva, 2006*
A.
capayaensis González-Sponga, 2006*
A.
caracensis González-Sponga, 2006*
A.
catuaroi González-Sponga, 2006*
A.
caucaguitensis González-Sponga, 2006*
A. chagasi Giupponi, Vasconcelos & Lourenço, 2009*
A.
charlescorfieldi Lourenço, 2001*
A.
chirimakei González-Sponga, 2006*
A.
claviformis González-Sponga, 2006*
A.
coinaui Lourenço, 1982
A. columbiana Lourenço, 1991
A. cryptozoicus Lourenço, 2005*
A.
cumbensis González-Sponga, 2006*
A.
curariensis González-Sponga, 2006*
A. cussinii Borelli,
1910
A. dekeyseri Lourenço, 1982
A.
deniseae Lourenço, 1997
A. diegorojasi Rojas-Runjaic, 2005*
A. dorae Botero-Trujillo, 2008*
A. ehrlichi Lourenço, 1994
A.
elguapoi González-Sponga, 2006*
A. elisabethae Lourenço, 2003*
A. evellynae Lourenço, 2004*
A. faguasi Botero-Trujillo, 2009*
A. feae (Borelli,
1911)
A. festae Borelli, 1899
A. franckei Lourenço, 1982
A. gorgonae Lourenço & Flórez, 1989
A.
guiripaensis González-Sponga, 2006*
A. guyanensis Lourenço & Monod, 1999*
A. hasshy Teruel & Roncallo, 2008*
A.
inoae González-Sponga, 2006*
A. kuryi Giupponi, Vasconcelos & Lourenço, 2009*
A. leilae Lourenço, 1999*
A. luciae Lourenço,
1984
A.
maniapurensis González-Sponga, 2006*
A. maranhensis Lourenço, 1987
A. mariaelenae Lourenço, 1999*
A. mariaterezae Lourenço, 1987
A. mauryi Lourenço, 1982
A.
meridana González-Sponga, 2006*
A. myriamae Botero-Trujillo, 2007*
A. nairae Lourenço, 2004*
A.
norae González-Sponga, 2006*
A.
paoensis González-Sponga, 2006*
A.
paracotoensis González-Sponga, 2006*
A.
plataensis González-Sponga, 2006*
A. platnicki Lourenço,
1993
A.
principalis González-Sponga, 2006*
A. pydanieli Lourenço, 1982
A.
riocaurensis González-Sponga, 2006*
A. riochicoi González-Sponga, 2006*
A. riomachensis Rojas-Runjaic, Portillo-Quintero & Borges, 2008*
A. sabineae Lourenço, 2001*
A.
sanchezi González-Sponga, 2006*
A.
sepulvedai González-Sponga, 2006*
A.
singularis González-Sponga, 2006*
A. terueli Kovarík, 2006*
A. tolimana Teruel & Garcia, 2007*
A. turumbanensis Gonzáles-Sponga, 1990
A.
venezuelensis Gonzáles-Sponga, 1972
A. volschenki Botero-Trujillo, 2009*
A.
zuliana González-Sponga, 2006*
|
| |
|
Androctonus
Ehrenberg, 1828
A. afghanus Lourenço & Qi, 2006*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi (Audouin,1826)
A. australis (Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. bicolor Ehrenberg, 1828
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Yhtier & Leguin, 2009*
A. sergenti Vachon, 1948
A. togolensis Lourenço, 2008*
|
Anomalobuthus
Kraepelin, 1900
A. rickmersi Kraepelin,
1900
|
Apistobuthus
Finnegan, 1932
A. pterygocercus Finnegan,
1932
A. susanae Lourenço, 1998
|
Australobuthus
Locket, 1990 A. xerolimniorum Locket,
1990
|
Babycurus Karsch,
1886
B. ansorgei Hirst, 1911
B. buettneri Karsch, 1886
B. centrurimorphus Karsch, 1886*
B. exquisitus Lowe, 2000*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. ornatus Werner, 1936
B. pictus Pocock, 1896*
B. solegladi Lourenço, 2005*
B. somalicus Hirst, 1907
B. subpunctatus Borelli, 1925
B. ugartei Kovarik, 2000*
B. wituensis Kraepelin, 1913
B. zambonellii Borelli, 1902
|
Baloorthochirus
Kovarik, 1996
B. becvari Kovarík,
1996
|
Birulatus Vachon,
1974
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
|
Buthacus Birula,
1908
B. arenicola (Simon, 1885)
B. birulai Lourenço, 2006*
B. buettikeri Hendrixson, 2006*
B. calviceps (Pocock, 1900)
B. clevai Lourenço, 2001*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (Nomen dubium)
B. huberi Lourenço, 2001*
B. leptochelys (Ehrenberg, 1829)
B. macrocentrus (Ehrenberg, 1828)*
B. mahraouii Lourenço, 2004*
B. maliensis Lourenço & Qi, 2007*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. occidentalis Lourenço, 2000*
B. pakistanensis Lourenço & Qi, 2006*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. willamsi Lourenço & Leguin, 2009*
B. ziegleri Lourenço, 2000*
|
Butheoloides
Hirst, 1925
B. annieae Lourenço,
1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. hirsti Lourenço, 1996
B.
maroccanus Hirst, 1925
B. milloti
Vachon, 1948
B. monodi Vachon, 1950
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. schwendingeri Lourenço, 2002*
B. wilsoni Lourenço, 1995
|
Butheolus Simon,
1882
B. andersoni (Pocock, 1895)*
B. anthracinus (Pocock, 1895)
B. arabicus Lourenço & Qi, 2006*
B.
ferrugineus Kraepelin, 1898
B.
gallagheri Vachon, 1980
B. thalassinus
Simon, 1882
B. villosus Hendrixson, 2006*
|
Buthiscus Birula,
1905
B. bicalcaratus Birula, 1905
|
Buthoscorpio Werner, 1936
B. jinnahii Amir, Kamaluddin & Jabbar, 2005*!
B. politus (Pocock, 1899)
B. rahmatii Amir, Kamaluddin & Jabbar, 2005*!
B. sarasinorum (Karsch, 1891)
! B. jinnahii & B. rahmatii was described in the non-valid genus
Stenochirus by the authors. I have placed them in Buthoscorpio, which is
the valid name for Stenochirus, but formally this has to be done in a scientific journal before it is valid. |
Buthus Leach,
1815
B. albengai Lourenço, 2003*
B. atlantis Pocock, 1889
B. barcaeus Birula, 1909*
B. barbouri Werner, 1932
B. berberensis Pocock, 1900*
B. bonito Lourenço & Geniez, 2005*
B. brignolii Lourenço, 2003*
B. chambiensis Kovarik, 2006*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. ibericus Lourenço & Vachon, 2004*
B. insolitus Borelli, 1925
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. jianxinae Lourenço, 2005*
B. lienhardi Lourenço, 2003*
B. malhommei Vachon, 1949*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. paris (C. L. Koch, 1839)*
B. rochati Lourenço, 2003*
B. tassili Lourenço, 2002*
B. tunetanus (Herbst, 1800)*
B. yemenensis Lourenço, 2008*
|
Centruroides Marx,
1890
C. alayoni Armas, 1999*
C.
anchorellus Armas, 1976
C.
arctimanus (Armas, 1976)
C. baergi Hoffmann, 1932*
C. balsasensis Ponce & Francke, 2004*
C. bani
Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis
(Pocock, 1898)
C. bertholdii (Thorell,
1876)
C.
bicolor (Pocock, 1898)
C.
chamulaensis Hoffmann, 1932
C.
chiapanensis Hoffmann, 1932
C. elegans
(Thorell, 1876)
C.
exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul
(Meise, 1934)
C. farri Armas, 1976
C.
flavopictus (Pocock, 1898)
C. fulvipes
(Pocock, 1898)
C. galano Teruel, 2001*
C.
gracilis (Latreille, 1804)
C.
griseus (C. L. Koch, 1844)
C.
guanensis Franganillo, 1930
C. hentzi (Banks, 1900)
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. mahnerti Lourenço, 1983
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mariaorum Santiago-Blay, 2009*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. morenoi Mello-Leitao, 1945*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus
(Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. platnicki
Armas, 1981
C. pococki Sissom & Francke,
1983
C. polito Teruel, 2007*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. subgranosus (Kraepelin, 1898)
C. suffusus (Pocock, 1902)
C. tecomanus Hoffmann, 1932*
C. testaceus (DeGeer, 1778)
C. thorellii (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. vittatus (Say, 1821)
|
Charmus Karsch,
1879
C. brignolii Lourenço, 2000*
C. indicus Hirst, 1915
C. laneus Karsch, 1879
C. minor Lourenço, 2002*
C. sinhagadensis
Tikader & Bastawade, 1983
|
|
Cicileiurus Teruel,
2007*
C. monticola Teruel, 2007*
|
Cicileus Vachon,
1948
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
|
Compsobuthus
Vachon, 1949
C. abyssinicus (Birula,
1903)
C. acutecarinatus (Simon,
1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov,
1973
C. becvari Kovarik, 2003*
C. berlandi Vachon, 1950
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. fuscatus Hendrixson, 2006*
C. garyi Lourenço & Vachon, 2001*
C. humaae Amir, Kamaluddin & Kahn, 2005*
C. jakesi Kovarik, 2003*
C. jordanensis
Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kafkai Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. klaptoczi (Birula, 1909)
C. longipalpis Levy, Amitai
& Shulov, 1973
C. maindroni (Kraepelin,
1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. polisi
Lowe, 2001*
C. plutenkoi Kovarik, 2003*
C. rugosulus (Pocock,
1900)
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. vachoni Sisson, 1994
C.
werneri (Birula, 1908)
C. williamsi Lourenço, 1999*
|
Congobuthus Lourenço,
1999*
C. fagei Lourenço, 1999*
|
Darchenia Vachon,
1977 D. bernadettae Vachon, 1977
|
Egyptobuthus Lourenço,
1999*
E. vaissadei Lourenço, 1999*
|
Grosphus Simon,
1880
G. ankarafantsika Lourenço, 2003*
G. ankarana Lourenço & Goodman, 2003*
G. annulatus Fage, 1929
G.
bistriatus Kraepelin, 1900
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. feti
Lourenço, 1996
G. flavopiceus Kraepelin,
1900
G. garciai Lourenço, 2001*
G. goudoti Lourenço & Goodman, 2006*
G. grandidieri Kraepelin,
1900
G. hirtus Kraepelin, 1900
G. intertidalis Lourenço, 1999*
G.
limbatus (Pocock, 1889)
G.
madagascariensis (Gervais, 1843)
G. mandena Lourenço, 2005*
G. mahafaliensis Lourenço, Goodman & Ramilijaona, 2004*
G. mayottensis Lourenço & Goodman, 2009*
G. olgae Lourenço, 2004*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
|
Hemibuthus Pocock, 1900
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
|
Hemilychas Hirst,
1911
H. alexandrinus (Hirst, 1911)
|
Himalayotityobuthus
Lourenço, 1997
H. alejandrae Lourenço, 2003*
H. martensi Lourenço,
1997
|
Hottentotta
Birula, 1908
H. alticola (Pocock, 1895)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004*
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. franzwerneri (Birula, 1914)
H. gentili (Pallary, 1924)*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1975)
H. niloticus (Birula, 1928)*
H. pachyurus (Pocock, 1897)*
H. penjabensis Kovarik, 2007*
H. polystictus (Pocock, 1896)
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1828)
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. socotrensis (Pocock, 1889)
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (Nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trilineatus (Peters, 1861)
H.
zagrosensis Kovarík, 1997
|
Iranobuthus
Kovarík, 1997
I. krali Kovarík,
1997
|
Isometroides
Keyserling, 1885
I. vescus (Karsch,
1880)
|
|
Isometrus Ehrenberg,
1828
I. acanthurus Pocock, 1899
I.
assamensis Oates, 1888
I. basilicus
Karsch, 1879
I. besucheti Vachon,
1982
I. brachycentrus Pocock, 1899
I.
corbeti Tikander & Batawade, 1983
I. bilyi Kovarík, 2003*
I.
formosus Pocock, 1894
I. garyi Lourenço & Huber, 2002*
I. hainanensis Lourenço, 2005*
I. heimi Vachon, 1976
I. isadensis Tikander & Bastawade,
1983
I. khammamensis Kovarík, 2003*
I. krasenskyi Kovarík, 1998
I.
kurkai Kovarík, 1997
I. loebli
(Vachon, 1982)*
I. maculatus
(DeGeer, 1778)
I. melanodactylus (L. Koch,
1867)
I. navaiae Kovarík, 1998
I. petrzelkai Kovarík, 2003*
I. problematicus Kovarík, 2003*
I.
rigidulus Pocock, 1897
I. sankeriensis
Tikander & Bastawade, 1983
I. thurstoni
Pocock, 1893
I. thwaitesi Pocock,
1897
I. tibetanus Lourenço & Zhu, 2008*
I. vittatus Pocock, 1900
I.
zideki Kovarík, 1994
|
Karasbergia
Hewitt, 1913
K. methueni Hewitt, 1913 |
Kraepelinia
Vachon, 1974
K. palpator (Birula, 1903) |
Lanzatus Kovarik,
2001*
L. somalicus Kovarik, 2001* |
Leiurus Ehrenberg,
1828
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. nasheri Kovarik, 2007*
L. quinquestriatus (Ehrenberg, 1828)
L. savanicola Lourenço, Qi & Cloudsley-Thompson, 2006*
| |
Liobuthus Birula,
1898
L. kessleri Birula, 1898 |
Lissothus Vachon,
1948
L. bernardi Vachon, 1948
L.
occidentalis Vachon, 1950 |
Lychas C.L. Koch,
1845
L. albimanus Henderson, 1919
L.
asper (Pocock, 1891)
L. biharensis
Tikader & Bastawade, 1983
L. braueri
(Kraepelin, 1986)
L. buchardi Kovarík,
1997
L. burdoi (Simon, 1882)
L. ceylonensis Lourenço & Huber, 1999*
L.
farkasi Kovarik, 1997
L. feae (Thorell,
1889)
L. flavimanus (Thorell,
1888)
L. gravelyi Henderson, 1913
L.
hendersoni (Pocock, 1897)
L.
heurtaultae Kovarík, 1997
L. hillyardi
Kovarík, 1997
L. hosei (Pocock,
1891)
L. infuscatus (Pocock, 1891)
L.
kamshetensis Tikader & Bastawade,
1983
L. kharpadi Bastawade, 1986
L.
krali Kovarík, 1995
L. laevifrons
Pocock, 1897
L. Lourençoi Kovarík,
1997
L.
marmoreus (C.L. Koch, 1844)
L.
mjobergi Kraepelin, 1916
L. mucronatus
(Fabricius. 1798)
L. nigristernis (Pocock,
1899)
L. obsti Kraepelin, 1913
L.
perfidus (Keyserling, 1885)
L. rackae
Kovarík, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. shoplandi (Oates, 1888)
L. srilankensis Lorenco, 1997
L. tricarinatus (Simon, 1884)
L. variatus (Thorell, 1876) |
|
Lychasioides
Vachon, 1974
L. amieti Vachon, 1974 |
Mauritanobuthus Qi & Lourenço, 2007*
M. geniezi Qi &Lourenço, 2007*
|
Mesobuthus Vachon,
1950
M. agnetis (Werner, 1936)
M. caucasicus (Nordmann,
1840)
M. cyprius Gantenbein & Kropf, 2000*
M.
eupeus (C.L. Koch, 1839)
M. extremus
(Werner, 1936)
M. gibbosus (Brulli,
1832)
M.
macmahoni (Pocock, 1900)
M. martensii
(Karsch, 1879)
M. nigrocinctus (Ehrenberg,
1828)*
M. songi Lourenço, Qi & Zhu, 2005*
M. vesiculatus (Pocock,
1899)
|
|
Mesotityus
Gonzalez-Sponga, 1981
M. vondangeli
González-Sponga, 1981 |
Microananteris
Lourenço, 2003*
M. minor Lourenço, 2003* |
Microbuthus
Kraepelin, 1898
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
M. pusillus Kraepelin, 1898 |
Microcharmus Lourenço, 1995
M. bemaraha Lourenço, Goodman & Fisher, 2006*
M. cloudsleythompsoni Lourenço, 1995
M. confluenciatus Lourenço, Goodman & Fisher, 2006*
M. duhemi Lourenço, Goodman & Fisher, 2006*
M. fisheri Lourenço, 1998
M. hauseri Lourenço, 1996
M. jussarae Lourenço, 1996*
M. maculatus Lourenço, 1996
M. madagascariensis Lourenço, 1999*
M. pauliani (Lourenço, 2004)*
M. sabineae Lourenço, 1996
M. variegatus Lourenço, Goodman & Fisher, 2006*
M. violaceous Lourenço, Goodman & Fisher, 2006*
|
Microtityus
Kjellesvig-Waering, 1966
M. angelaerrosae Gonzáles-Sponga, 2001*
M. biordi González-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. capayaensis González-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae González-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. farleyi Teruel, 2000*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M.
jaumei Armas, 1974
M. joseantonioi González-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis González-Sponga, 2001*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. rickyi Kjellesvig-Waering, 1966
M. sevciki González-Sponga, 2001*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. virginiae Armas, 1999*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus González-Sponga, 2001*
|
Neobuthus Hirst,
1911
N. cloudsleythompsoni Lourenço, 2001*
N. berberensis (Hirst, 1911)
N. sudanensis Lourenço, 2005*
|
Neogrosphus
Lourenço, 1995
N. blanci Lourenço,
1996
N. griveaudi (Vachon, 1969) |
Neoprotobuthus
Lourenço, 2000*
N. intermedius Lourenço,
2000* |
Odontobuthus
Vachon, 1950
O. bidentatus (Lourenço & Pezier, 2002)*
O. doriae (Thorell,
1876)
O. odonturus (Pocock, 1897)
|
Odonturus Karsch,
1879
O.
dentatus Karsch, 1879 |
Orthochiroides
Kovarík, 1998
O. insularis (Pocock, 1889)*
O. socotrensis Kovarík, 2004*
O. vachoni Kovarík, 1998
|
Orthochirus
Karsch, 1891
O. afghanus Kovarik, 2004*
O. aristidis (Simon, 1882)
O. bicolor (Pocock, 1897)
O. blandini (Lourenço & Vachon, 1997)*
O. danielleae (Lourenço & Vachon, 1997)*
O. erardi (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fuscipes (Pocock, 1900)
O. glabrifrons(Kraepelin, 1903)*
O. goyffoni (Lourenço & Vachon, 1995)*
O. gromovi Kovarik, 2004*
O. gruberi Kovarik & Fet, 2006*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. kaspareki (Lourenço & Huber, 2000)*
O. kinzelbachi (Lourenço & Huber, 2000)*
O. krishnai Tikader & Bastawade, 1983
O. monodi (Lourenço & Vachon, 1997)*
O. pallidus (Pocock, 1897)
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. sobotniki Kovarik, 2004*
O. stockwelli (Lourenço & Vachon, 1995)*
O. varius Kovarik, 2004*
O. zagrosensis Kovarik, 2004*
|
Pantobuthus
Lourenço & Duhem, 2009*
P. complicatus Lourenço & Duhem, 2009*
|
Parabuthus Pocock,
1890
P. brevimanus (Thorell, 1876)
P.
calvus Purcell, 1898
P. capensis
(Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral,
1980
P. eritreaensis Kovarik, 2003*
P. gracilis Lamoral, 1979
P.
granimanus Pocock, 1895
P.
granulatus (Ehrenberg, 1831)
P.
heterurus Pocock, 1899
P. hunteri
Pocock, 1895
P. kalaharicus Lamoral,
1977
P. kraepelini Werner, 1902
P.
kuanyamarum Monrad, 1937
P. laevifrons
(Simon, 1888)
P.
liosoma (Ehrenberg, 1828)
P.
maximus Werner, 1913
P. mossambicensis (Peters,
1861)
P. muelleri Prendini, 2000*
P.
namibensis Lamoral, 1979
P. nanus
Lamoral, 1979
P.
pallidus Pocock, 1895
P. planicauda
(Pocock, 1889)
P. raudus (Simon,
1888)
P. schlechteri Purcell, 1899
P.
stridulus Hewitt, 1913
P.
transvaalicus Purcell, 1899
P.
villosus (Peters, 1862)
| |
Pectinibuthus Fet,
1984
P. birulai Fet, 1984 |
Physoctonus
Mello-Leitao, 1934*
P. debilis (C. L. Koch, 1840)*
|
Plesiobuthus
Pocock, 1900
P. paradoxus Pocock, 1900
|
Polisius Fet,
Capes & Sissom, 2001*
P. persicus Fet,
Capes & Sissom, 2001* |
Psammobuthus
Birula, 1911
P. zarudnyi Birula, 1911
|
Pseudolissothus
Lourenço, 2001*
P. pussilus Lourenço, 2001*
|
Pseudolychas
Kraepelin, 1911
P. ochraceus (Hirst,
1911)
P. pegleri (Purcell, 1901)
P.
transvaalicus Lawrence, 1961
|
|
Pseudouroplectes
Lourenço, 1995
P. betschi Lourenço, 1995
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999* |
Razianus
Farzanpay, 1987
R. zarudnyi (Birula, 1903)
|
Rhopalurus
Thorell, 1876
R. abudi Armas & Marcano
Fondeur, 1987
R. acromelas Lutz & Mello,
1922
R. agamemmon (C. L. Koch,
1839)
R. amazonicus Lourenço, 1986
R. bonettii Armas 1999*
R. caribensis Teruel & Roncallo, 2008*
R. crassicauda Caporiacco, 1947*
R.
garridoi Armas, 1974
R. guanambiensis
Lenarducci, Pinto-da-Rocha & Lucas, 2005*
R. gibarae Teruel, 2006*
R. granulimanus Teruel, 2006*
R.
junceus (Herbst, 1800)
R. lacrau
Lourenço & Pinto-da-Rocha, 1997
R.
laticauda Thorell, 1876
R. melloleitaoi Teruel & Armas, 2006*
R. pintoi Mello-Leităo, 1932*
R.
princeps (Karsch, 1879)
R. rochai Borelli, 1910
R. virkkii Santiago-Blay, 2009*
|
Sabinebuthus
Lourenço, 2001*
S. elegans Lourenço, 2001*
|
Saharobuthus
Lourenço & Duhem, 2009*
S. elegans Lourenço & Duhem, 2009*
|
|
Sassandiothus
Farzanpay, 1987
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900)
|
Somalibuthus
Kovarík, 1998
S. demisi Kovarík, 1998 |
Somalicharmus
Kovarík, 1998
S. withmanae Kovarík, 1998 |
Thaicharmus
Kovarík, 1995
T. lowei Kovarík, Soleglad & Fet, 2007*
T. mahunkai Kovarík, 1995
|
Tityobuthus
Pocock, 1893
T. antsingy Lourenço & Goodman, 2004*
T. baroni (Pocock, 1890)
T. betschi Lourenço, Qi & Goodman, 2008*
T. chelbergorum Lourenço, Qi & Goodman, 2008*
T. darainensis Lourenço & Goodman, 2002*
T. dastychi Lourenço, 1997
T. griswoldi Lourenço, 2000*
T. guillaumeti
Lourenço, 1995
T. ivohibe Lourenço & Goodman, 1999*
T. judsoni Lourenço,
1996
T. manonae Lourenço, 2000*
T. mccarteri Lourenço, Qi & Goodman, 2008*
T. monodi Lourenço, 2000*
T. pallidus Lourenço, 2004*
T.
parrilloi Lourenço, 1996
T. petrae
Lourenço, 1996
T. pococki Lourenço, 1995
T. rakotondravonyi Lourenço, 2003*
|
Tityopsis Armas,
1974
T. aliciae Armas & Frias, 1998*
T. inaequalis (Armas, 1974)
T.
inexpectata (Moreno, 1940) |
Tityus C.L. Koch,
1836
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. abudi Armas, 1999*
T. acananensis González-Sponga, 2009*
T. adisi Lourenço, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi González-Sponga, 2001*
T. altithronus Armas, 1999*
T. anasilviae Armas & Abud Antun, 2004*
T. androcottoides (Karsch, 1879)
T. anduzei González-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. antioquensis Lourenço& Otero Patińo, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai González-Sponga, 1985
T. argentinus Borelli,
1899
T. asthenes Pocock, 1893
T.
atriventer Pocock, 1897
T. bahiensis
(Perty, 1833)
T. bahoruco Teruel & Armas, 2006*
T. barquisimetanus
González-Sponga, 1994
T. bastosi Lourenço,
1984
T. bellulus Armas, 1999*
T.
betschi Lourenço, 1992
T. birabeni
Abalos, 1955
T. blanci Lourenço,
1994
T. blaseri Mello-Leităo, 1931
T. boconoensis González-Sponga, 1981
T.
bolivanus Kraepelin, 1895
T. brazilae
Lourenço & Eickstedt, 1984
T. breweri
González-Sponga, 1997
T. cachipalensis González-Sponga, 2002*
T. caesarbarrioi González-Sponga, 2001*
T. canopensis Lourenço, 2002*
T. carabobensis González-Sponga, 1987
T. carinatoides Mello-Leităo, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carvalhoi Mello-Leităo, 1945
T. cerroazul Lourenço,
1986
T. charalaensis Mello-Leităo,
1940
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. clathratus C. L. Koch, 1844
T.
columbianus (Thorell, 1876)
T.
confluens Borelli, 1899
T. costatus
(Karsch, 1879)
T. crassimanus (Thorell,
1876)
T. cuellari Lourenço, 1994
T.
culebrensis González-Sponga, 1994
T. cylindricus (Karsch, 1879)*
T.
dasyurus Pocock, 1897
T. dedoslargos
Francke & Stockwell, 1987
T. demangei
Lourenço, 1981
T. dinizi Lourenço,
1997
T. discrepans (Karsch, 1879)
T. dorae González-Sponga, 2001*
T. duacaensis González-Sponga, 2007*
T. dulceae González-Sponga, 2006*
T. dupouyi González-Sponga, 1987
T.
ebanoverde Armas, 1999*
T. ecuadorensis
Kraepelin, 1896
T. elii (Armas & Marcano
Fondeur, 1992)*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi González-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. erikae Lourenço, 1999*
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leităo, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessôa, 1935
T. festae
Borelli, 1899
T. filodendron
González-Sponga, 1981
T. florezi Lourenço,
2000*
T. footei Chamberlin, 1916
T.
forcipula (Gervais, 1843)
T. fuhrmanni Kraepelin, 1914
T. funestus Hirst,
1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. guaricoensis Gonzalez-Sponga, 2004*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus
Mello-Leităo, 1934
T. insignis (Pocock,
1889)
T. intermedius Borelli, 1899
T. irapaensis González-Sponga, 2002*
T.
isabelceciliae González-Sponga, D'Suze &
Sevcik, 2001*
T. ivani González-Sponga, 2008*
T. ivicnancor González-Sponga,
1997
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai González-Sponga, 2007*
T. kuryi Lourenço, 1997
T. lancinii González-Sponga, 1972
T. lokiae Lourenço, 2005*
T. longidigitus González-Sponga, 2008*
T. Lourençoi Flórez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis González-Sponga, 2007*
T. manakai González-Sponga, 2004*
T. maniapurensis González-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. martinpaechi Lourenço, 2001*
T. matthieseni Lourenço & Pinto-da-Rocha, 2000
T. mattogrossensis Borelli,
1901
T. maturinensis González-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis González-Sponga, 1974
T. mongei Lourenço, 1996
T. mraceki Kovarik, 2007*
T. mucusunamensis González-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leităo, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leităo, 1940
T. neoespartanus González-Sponga, 1996
T. nororientalis González-Sponga, 1996
T. obispoi González-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus González-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis González-Sponga, 2007*
T. paraguayensis Kraepelin, 1895
T.
parvulus Kraepelin, 1914
T. paulistorum Lourenço & Qi, 2006*
T.
perijanensis González-Sponga, 1994
T.
pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri
González-Sponga, 1981
T. pococki Hirst,
1907
T. portoplatensis Armas & Marcano
Fondeur, 1992
T. potameis Lourenço & Leăo Giupponi, 2004*
T. prancei Lourenço,
2000*
T. proseni Abalos, 1954
T.
pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis González-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus (Armas, 1982)*
T. ramirezi Esquivel de Verde, 1969 [nomen dubium]
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis González-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi González-Sponga, 1996
T. romeroi González-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rufofuscus Pocock, 1897
T. rugosus Schenkel, 1932
T. rusmelyae González-Sponga, D'Suze & Sevcik, 2001*
T. sabinae Lourenço, 1994
T. sanarensis González-Sponga, 1997
T. sanfernandoi González-Sponga, 2008*
T. sarisarinamensis González-Sponga, 2002*
T. sastrei Lourenço
& Flórez, 1990
T. septentrionalis Armas
& Abud Antun, 2004*
T. serrulatus Lutz &
Mello, 1922
T. shiriana González-Sponga,
1991
T. silvestris Pocock, 1897
T.
simonsi Pocock, 1900
T. soratensis
Kraepelin, 1912
T. stigmurus (Thorell,
1876)
T. strandi Werner, 1939
T. surimeridensis González-Sponga, 2002*
T.
surorientalis González-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi González-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. thelyacanthus Mello-Leităo, 1933
T. trinitatis Pocock,
1897
T. trivittatus Kraepelin,
1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leităo, 1931
T. unus Lourenço & Pinto-da-Rocha, 2000
T. uquirensis González-Sponga, 2001*
T. urachichensis González-Sponga, 2007*
T. urbinai
Scorza, 1952
T. uruguayensis Borelli,
1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis González-Sponga, 1981
T. ventuarensis González-Sponga, 2009*
T. walli González-Sponga & Wall Gonzalez, 2007*
T. wayuu Rojas-Runjaic & Armas, 2007*
T. yerenai González-Sponga, 2009*
T. zulianus González-Sponga, 1981
|
|
Troglorhopalurus Lourenço, Baptista & Giupponi,
2004*
T. translucidus Lourenço, Baptista & Giupponi,
2004*
| Troglotityobuthus Lourenço,
2000*
T. gracilis (Fage, 1946)
|
Uroplectes Peters,
1861
U. fischeri (Karsch, 1879)
U.
andreae Pocock, 1889
U. carinatus
(Pocock, 1890)
U. chubbi Hirst,
1911
U. fischeri (Karsch, 1879)
U.
flavoviridis Peters, 1861
U. formosus
Pocock, 1890
U. gracilior Hewitt,
1914
U. insignis Pocock, 1890
U.
lineatus (C. L. Koch, 1844)
U.
longimanus Werner, 1936
U. marlothi
Purcell, 1901
U. ngangelarum Monard,
1930
U. occidentalis Simon, 1876
U.
olivaceus Pocock, 1896
U.
otjimbinguensis (Karsch, 1879)
U.
pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner,
1913
U. pilosus (Thorell, 1876)
U.
planimanus (Karsch, 1879)
U.
schlechteri Purcell, 1901
U. schubotzi
Kraepelin, 1929
U. silvestrii Borelli,
1913
U. teretipes Lawrence, 1966
U.
triangulifer (Thorell, 1876)
U.
tumidimanus Lamoral, 1979
U. variegatus
(C. L. Koch, 1844)
U. vittatus (Thorell,
1876)
U. xanthogrammus Pocock,
1897
| |
Uroplectoides
Lourenço, 1998
U. abyssinicus Lourenço,
1998
U. emiliae (Werner, 1916) |
Vachoniolus Levy,
Amitai & Shulov, 1973
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
|
Vachonus Tikader
& Bastawade, 1983
V. astrostriatus
(Pocock, 1897)
V. rajasthanicus Tikader
& Bastawade, 1983
|
Zabius Thorell, 1893
Z. birabeni Mello-Laeităo, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*
|
|
|
