| With 89 genera (one extinct) and 990 species [Updated 25.04.13],
this family is the largest of the scorpion families. This
family is widespread around the world (not found in Antarctica
and New Zealand), and are found in tropical, subtropical and
partly in temperate habitats.
Volschenk, Mattoni & Prendini (2008) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus and Neoprotobuthus) and 15 species are all transfered to Buthidae. Information about the old family Microcharmidae.
Kovarik (2007) has published a revision of the genus Hottentotta. Note that
the medical important Mesobuthus tamulus and two other Asian Mesobuthus have been
transfered to Hottentotta. Kovarik's revision can be
downloaded HERE (browse down to issue 58).
Paraorthochirus has been synonymized with Orthochirus by Navidpour et al., 2008).
The members of this family are small and medium sized
scorpions with a triangular sternum (some genera have a more
pentagonal sternum). In many genera (e.g., Androctonus,
Apistobuthus, Parabuthus, and other Old World
genera) the cauda is usually strong and powerful and the
pedipalps are often very slender whereas other genera have
elongate appendages, especially the males (e.g.,
Centruroides, Lychas, Isometrus and
related genera). Many species are yellow or brown (or
variations of these colors), but black forms are also present.
Some species have quite intricate and attractive patterns
(e.g., Lychas and Isometrus) and colours (e.g.,
Centruroides and Uroplectes). Sizes range from
about 20 mm (like Microtityus and Microbuthus)
to over 120 mm (as in Androctonus, some
Centruroides, Apistobuthus and others).
An indentification key to the genus Compsobuthus was published by Kovarik (2012) [Free full text].
Several species in this family are highly toxic, but fewer
than 20 can be lethal to man. The most notorious species are
found in the genus Androctonus, Centruroides, Hottentotta,
Leiurus, Parabuthus and Tityus. See Scorpions
of medical importance page for further information.
Unlike most other scorpion families, the Buthidae are of
both scientific and public interest due to their medical
importance in many parts of the world. Several species have
also made their way into the hobby. Many of these are safe to
be kept by experienced collectors, but the most dangerous
species should be avoided.
Androctonus crassicauda habitat picture from Kuwait are located HERE.
Photo of Parabuthus leiosoma (left)
by Jan Ove Rein (C)
Photo of Hottentotta sp.
(right) by Jan Ove Rein (C)
SPECIES
FILES:
This list of genera and
species is based on Fet
et al.(2000). Subfamilies and subspecies is not included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000) |
Afghanobuthus
Lourenço, 2005*
A. naumanni Lourenço, 2005*
|
|
Afroisometrus
Kovarík, 1997
A. minshullae (Fitzpatrick,
1994)
|
Akentrobuthus
Lamoral, 1976
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976 |
Alayotityus Armas,
1973
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A.
juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus
Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas,
1973
|
Ananteris Thorell,
1891
A. arcadioi Botero-Trujillo, 2008*
A. ashmolei Lourenço, 1981
A. asuncionensis González-Sponga, 2006*
A. balzanii Thorell, 1891
A. barinensis González-Sponga, 2006*
A. bernabei Giupponi, Vasconcelos & Lourenço, 2009*
A. bianchinii Lourenço, Aguiar-Neto & Limeira-de-Oliveira, 2009*
A. bonito Lourenço, 2012*
A. cachimboensis Lourenço, Motta & da Silva, 2006*
A.
capayaensis González-Sponga, 2006*
A.
caracensis González-Sponga, 2006*
A.
catuaroi González-Sponga, 2006*
A.
caucaguitensis González-Sponga, 2006*
A. chagasi Giupponi, Vasconcelos & Lourenço, 2009*
A.
charlescorfieldi Lourenço, 2001*
A.
chirimakei González-Sponga, 2006*
A.
claviformis González-Sponga, 2006*
A.
coinaui Lourenço, 1982
A. columbiana Lourenço, 1991
A. cryptozoicus Lourenço, 2005*
A. cumbensis González-Sponga, 2006*
A. curariensis González-Sponga, 2006*
A. cussinii Borelli, 1910
A. dekeyseri Lourenço, 1982
A. deniseae Lourenço, 1997
A. diegorojasi Rojas-Runjaic, 2005*
A. dorae Botero-Trujillo, 2008*
A. ehrlichi Lourenço, 1994
A. elguapoi González-Sponga, 2006*
A. elisabethae Lourenço, 2003*
A. evellynae Lourenço, 2004*
A. faguasi Botero-Trujillo, 2009*
A. festae Borelli, 1899
A. franckei Lourenço, 1982
A. gorgonae Lourenço & Flórez, 1989
A.
guiripaensis González-Sponga, 2006*
A. guyanensis Lourenço & Monod, 1999*
A. inoae González-Sponga, 2006*
A. intermedia Lourenço, 2012*
A. kuryi Giupponi, Vasconcelos & Lourenço, 2009*
A. leilae Lourenço, 1999*
A. luciae Lourenço, 1984
A. madeirensis Lourenço & Duhem, 2010*
A. maniapurensis González-Sponga, 2006*
A. maranhensis Lourenço, 1987
A. mariaelenae Lourenço, 1999*
A. mariaterezae Lourenço, 1987
A. mauryi Lourenço, 1982
A. meridana González-Sponga, 2006*
A. myriamae Botero-Trujillo, 2007*
A. nairae Lourenço, 2004*
A. norae González-Sponga, 2006*
A. ochoai Botero-Trujillo & Florez D, 2011*
A. palmari Botero-Trujillo & Noriega, 2011*
A. paoensis González-Sponga, 2006*
A. paracotoensis González-Sponga, 2006*
A. plataensis González-Sponga, 2006*
A. platnicki Lourenço, 1993
A. principalis González-Sponga, 2006*
A. pydanieli Lourenço, 1982
A. riocaurensis González-Sponga, 2006*
A. riochicoi González-Sponga, 2006*
A. riomachensis Rojas-Runjaic, Portillo-Quintero & Borges, 2008*
A. roraima Lourenço & Duhem, 2010*
A. sabineae Lourenço, 2001*
A. sanchezi González-Sponga, 2006*
A. sepulvedai González-Sponga, 2006*
A. singularis González-Sponga, 2006*
A. solimariae Botero-Trujillo & Florez D, 2011*
A. surinamensis Lourenço, 2012*
A. terueli Kovarík, 2006*
A. tolimana Teruel & Garcia, 2007*
A. turumbanensis Gonzáles-Sponga, 1990
A.
venezuelensis Gonzáles-Sponga, 1972
A. volschenki Botero-Trujillo, 2009*
A.
zuliana González-Sponga, 2006*
|
Ananteroides
Borelli, 1911*
A. feae Borelli, 1911*
| |
Androctonus
Ehrenberg, 1828
A. afghanus Lourenço & Qi, 2006*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi (Audouin,1826)
A. australis (Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. bicolor Ehrenberg, 1828
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Yhtier & Leguin, 2009*
A. pallidus Lourenço, Duhem & Cloudsley-Thompson, 2012*
A. sergenti Vachon, 1948
A. togolensis Lourenço, 2008*
|
Anomalobuthus
Kraepelin, 1900
A. rickmersi Kraepelin,
1900
|
Apistobuthus
Finnegan, 1932
A. pterygocercus Finnegan,
1932
A. susanae Lourenço, 1998
|
Australobuthus
Locket, 1990 A. xerolimniorum Locket,
1990
|
Babycurus Karsch,
1886
B. ansorgei Hirst, 1911
B. buettneri Karsch, 1886
B. centrurimorphus Karsch, 1886*
B. exquisitus Lowe, 2000*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. ornatus Werner, 1936
B. pictus Pocock, 1896*
B. solegladi Lourenço, 2005*
B. somalicus Hirst, 1907
B. subpunctatus Borelli, 1925
B. ugartei Kovarik, 2000*
B. wituensis Kraepelin, 1913
B. zambonellii Borelli, 1902
|
Baloorthochirus
Kovarik, 1996
B. becvari Kovarík,
1996
|
Birulatus Vachon,
1974
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
|
Buthacus Birula,
1908
B. agarwali Zambre & Lourenço, 2010*
B. arenicola (Simon, 1885)
B. birulai Lourenço, 2006*
B. buettikeri Hendrixson, 2006*
B. calviceps (Pocock, 1900)
B. clevai Lourenço, 2001*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (Nomen dubium)
B. golovatchi Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. huberi Lourenço, 2001*
B. leptochelys (Ehrenberg, 1829)
B. macrocentrus (Ehrenberg, 1828)*
B. mahraouii Lourenço, 2004*
B. maliensis Lourenço & Qi, 2007*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. occidentalis Lourenço, 2000*
B. pakistanensis Lourenço & Qi, 2006*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. willamsi Lourenço & Leguin, 2009*
B. ziegleri Lourenço, 2000*
|
Butheoloides
Hirst, 1925
B. annieae Lourenço,
1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. granulatus Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. hirsti Lourenço, 1996
B. littoralis Lourenço, Touloun & Boumezzough, 2011*
B. maroccanus Hirst, 1925
B. milloti Vachon, 1948
B. monodi Vachon, 1950
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. schwendingeri Lourenço, 2002*
B. slimanii Lourenço, 2010*
B. wilsoni Lourenço, 1995
|
Butheolus Simon,
1882
B. andersoni (Pocock, 1895)*
B. anthracinus (Pocock, 1895)
B. arabicus Lourenço & Qi, 2006*
B. gallagheri Vachon, 1980
B. pallidus Lourenço & Duhem, 2012*
B. thalassinus Simon, 1882
B. villosus Hendrixson, 2006*
|
Buthiscus Birula,
1905
B. bicalcaratus Birula, 1905
|
Buthoscorpio Werner, 1936
B. indicus Lourenço, 2012*
B. politus (Pocock, 1899)
B. rayalensis Javed, Rao, Mirza, Sanap & Tampal, 2010*
B. sarasinorum (Karsch, 1891)
NB! B. jinnahii Amir, Kamaluddin & Jabbar, 2005* and B. rahmatii Amir, Kamaluddin & Jabbar, 2005* was described in the non-valid genus Stenochirus by the authors. I have placed them in Buthoscorpio, which is
the valid name for Stenochirus, but formally this has to be done in a scientific journal before it is valid. Javed et al., 2010 have studied the descriptions of these two species and concluded that they do not belong to the genus Buthoscorpio. This decsision was also supported by Lourenco, 2012. The two taxa are now Buthidae incertae sedis and are not included in the total number species in Buthidae. |
Buthus Leach,
1815
B. albengai Lourenço, 2003*
B. amri Lourenço, Yagmur & Duhem 2010*
B. atlantis Pocock, 1889
B. awashensis Kovarik, 2011*
B. barcaeus Birula, 1909*
B. barbouri Werner, 1932
B. berberensis Pocock, 1900*
B. bonito Lourenço & Geniez, 2005*
B. boumalenii Touloun & Boumezzough, 2011*
B. brignolii Lourenço, 2003*
B. chambiensis Kovarik, 2006*
B. confluens Lourenço, Touloun & Boumezzough, 2012*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. egyptiensis Lourenço, 2012*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. elongatus Rossi, 2012*
B. hassanini Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. ibericus Lourenço & Vachon, 2004*
B. insolitus Borelli, 1925
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. israelis (Shulov & Amitai, 1959)*
B. jianxinae Lourenço, 2005*
B. kunti Yagmur, Koc & Lourenço, 2011*
B. lienhardi Lourenço, 2003*
B. malhommei Vachon, 1949*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. orientalis Lourenço & Simon, 2012*
B. paris (C. L. Koch, 1839)*
B. prudenti Lourenço & Leguin, 2012*
B. rochati Lourenço, 2003*
B. tassili Lourenço, 2002*
B. tunetanus (Herbst, 1800)*
B. yemenensis Lourenço, 2008*
|
Centruroides Marx,
1890
C. alayoni Armas, 1999*
C.
anchorellus Armas, 1976
C.
arctimanus (Armas, 1976)
C. baergi Hoffmann, 1932*
C. balsasensis Ponce & Francke, 2004*
C. bani
Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis (Pocock, 1898)
C. bertholdii (Thorell, 1876)
C. bicolor (Pocock, 1898)
C. chamela Ponce-Saavedra & Francke, 2011*
C. chamulaensis Hoffmann, 1932
C. chiapanensis Hoffmann, 1932
C. edwardsii (Gervais, 1843)*
C. elegans (Thorell, 1876)
C. exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul (Meise, 1934)
C. fallassisimus Armas & Trujillo, 2010*
C. farri Armas, 1976
C. flavopictus (Pocock, 1898)
C. fulvipes (Pocock, 1898)
C. galano Teruel, 2001*
C. gracilis (Latreille, 1804)
C. granosus (Thorell, 1876)*
C. griseus (C. L. Koch, 1844)
C. guanensis Franganillo, 1930
C. hentzi (Banks, 1900)
C. hirsuticauda Teruel, 2011*
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. mahnerti Lourenço, 1983
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mariaorum Santiago-Blay, 2009*
C. mascota Ponce-Saavedra & Francke, 2011*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus
(Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. platnicki
Armas, 1981
C. pococki Sissom & Francke,
1983
C. polito Teruel, 2007*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. sanandres Armas, Sarmiento & Florez 2012*
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. simplex (Thorell, 1876)*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. suffusus (Pocock, 1902)
C. tecomanus Hoffmann, 1932*
C. testaceus (DeGeer, 1778)
C. thorelli (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. villegasi Baldazo-Monsivaiz, Ponce-Saavedra & Flores-Moreno, 2013*
C. vittatus (Say, 1821)
|
Charmus Karsch,
1879
C. brignolii Lourenço, 2000*
C. indicus Hirst, 1915
C. laneus Karsch, 1879
C. minor Lourenço, 2002*
C. sinhagadensis
Tikader & Bastawade, 1983
|
|
Cicileiurus Teruel,
2007*
C. monticola Teruel, 2007*
|
Cicileus Vachon,
1948
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
|
Compsobuthus
Vachon, 1949
C. abyssinicus (Birula, 1903)
C. acutecarinatus (Simon, 1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov, 1973
C. atrostriatus (Pocock, 1897)*
C. becvari Kovarik, 2003*
C. berlandi Vachon, 1950
C. birulai Lourenço, Leguin & Duhem, 2010*
C. boucheti Lourenço, Duhem & Cloudsley-Thompson, 2012*
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. fuscatus Hendrixson, 2006*
C. garyi Lourenço & Vachon, 2001*
C. humaae Amir, Kamaluddin & Kahn, 2005*
C. jakesi Kovarik, 2003*
C. jordanensis
Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kafkai Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. klaptoczi (Birula, 1909)
C. krali Kovarik, 2012*
C. levyi Kovarik, 2012*
C. longipalpis Levy, Amitai & Shulov, 1973
C. lowei Lourenço & Duhem, 2012*
C. maindroni (Kraepelin, 1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. polisi
Lowe, 2001*
C. plutenkoi Kovarik, 2003*
C. rugosulus (Pocock,
1900)
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. sindicus Kovarik & Ahmed, 2011*
C. sobotniki Kovarik, 2004*
C. somalilandus Kovarik, 2012*
C. tassili Lourenço, 2010*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. vachoni Sisson, 1994
C. werneri (Birula, 1908)
C. williamsi Lourenço, 1999*
|
Congobuthus Lourenço,
1999*
C. fagei Lourenço, 1999*
|
Darchenia Vachon,
1977
D. bernadettae Vachon, 1977
|
Egyptobuthus Lourenço,
1999*
E. vaissadei Lourenço, 1999*
|
Grosphus Simon,
1880
G. ankarafantsika Lourenço, 2003*
G. ankarana Lourenço & Goodman, 2003*
G. annulatus Fage, 1929
G. bicolor Lourenço, 2012*
G. bistriatus Kraepelin, 1900
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. feti
Lourenço, 1996
G. flavopiceus Kraepelin,
1900
G. garciai Lourenço, 2001*
G. goudoti Lourenço & Goodman, 2006*
G. grandidieri Kraepelin,
1900
G. hirtus Kraepelin, 1900
G. intertidalis Lourenço, 1999*
G.
limbatus (Pocock, 1889)
G.
madagascariensis (Gervais, 1843)
G. mandena Lourenço, 2005*
G. mahafaliensis Lourenço, Goodman & Ramilijaona, 2004*
G. mayottensis Lourenço & Goodman, 2009*
G. olgae Lourenço, 2004*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
|
Femtobuthus Lowe, 2010*
F. shutuae Lowe, 2010*
|
Hemibuthus Pocock, 1900
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
|
|
Hemilychas Hirst,
1911
H. alexandrinus (Hirst, 1911)
|
Himalayotityobuthus
Lourenço, 1997
H. alejandrae Lourenço, 2003*
H. martensi Lourenço,
1997
|
Hottentotta
Birula, 1908
H. alticola (Pocock, 1895)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004*
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. flavidulus Teruel & Rein, 2010*
H. franzwerneri (Birula, 1914)
H. gentili (Pallary, 1924)*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarik & Kayedi, 2010*
H. mateui Lourenço, Duhem & Cloudsley-Thompson, 2012*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1975)
H. niloticus (Birula, 1928)*
H. pachyurus (Pocock, 1897)*
H. pellucidus Lowe, 2010*
H. penjabensis Kovarik, 2007*
H. polystictus (Pocock, 1896)
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1828)
H. saxinatans Lowe, 2010*
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. socotrensis (Pocock, 1889)
H. songi (Lourenço, Qi & Zhu, 2005)*
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (Nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trilineatus (Peters, 1861)
H.
zagrosensis Kovarík, 1997
|
Iranobuthus
Kovarík, 1997
I. krali Kovarík,
1997
|
Isometroides
Keyserling, 1885
I. vescus (Karsch,
1880)
|
|
Isometrus Ehrenberg,
1828
I. acanthurus Pocock, 1899
I. assamensis Oates, 1888
I. atherii Amir & Kamaluddin, 2008*
I. basilicus Karsch, 1879
I. besucheti Vachon, 1982
I. bilyi Kovarík, 2003*
I. brachycentrus Pocock, 1899
I. corbeti Tikander & Batawade, 1983
I. deharvengi Lourenço & Duhem, 2010*
I. formosus Pocock, 1894
I. garyi Lourenço & Huber, 2002*
I. hainanensis Lourenço, 2005*
I. heimi Vachon, 1976
I. isadensis Tikander & Bastawade, 1983
I. khammamensis Kovarík, 2003*
I. krasenskyi Kovarík, 1998
I. kurkai Kovarík, 1997
I. lao Lourenço & Leguin, 2012*
I. liaqatii Amir & Kamaluddin, 2008*
I. loebli (Vachon, 1982)*
I. maculatus (DeGeer, 1778)
I. melanodactylus (L. Koch, 1867)
I. navaiae Kovarík, 1998
I. petrzelkai Kovarík, 2003*
I. problematicus Kovarík, 2003*
I. rigidulus Pocock, 1897
I. thurstoni Pocock, 1893
I. thwaitesi Pocock, 1897
I. tibetanus Lourenço & Zhu, 2008*
I. vittatus Pocock, 1900
I. zideki Kovarík, 1994
|
Karasbergia
Hewitt, 1913
K. methueni Hewitt, 1913 |
Kraepelinia
Vachon, 1974
K. palpator (Birula, 1903) |
Lanzatus Kovarik,
2001*
L. somalicus Kovarik, 2001* |
Leiurus Ehrenberg,
1828
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. nasheri Kovarik, 2007*
L. quinquestriatus (Ehrenberg, 1828)
L. savanicola Lourenço, Qi & Cloudsley-Thompson, 2006*
| |
Liobuthus Birula,
1898
L. kessleri Birula, 1898 |
Lissothus Vachon,
1948
L. bernardi Vachon, 1948
L.
occidentalis Vachon, 1950 |
Lychas C.L. Koch,
1845
L. aareyensis Mirza & Sanap, 2010*
L. aberlenci Lourenço, 2013*
L. albimanus Henderson, 1919
L. asper (Pocock, 1891)
L. biharensis Tikader & Bastawade, 1983
L. braueri (Kraepelin, 1986)
L. buchardi Kovarík, 1997
L. burdoi (Simon, 1882)
L. ceylonensis Lourenço & Huber, 1999*
L. eliseanneae Lourenço, 2011*
L. farkasi Kovarik, 1997
L. feae (Thorell, 1889)
L. flavimanus (Thorell,
1888)
L. gravelyi Henderson, 1913
L. hendersoni (Pocock, 1897)
L. heurtaultae Kovarík, 1997
L. hillyardi Kovarík, 1997
L. hosei (Pocock, 1891)
L. inexpectatus Lourenço, 2011*
L. infuscatus (Pocock, 1891)
L. kaimana Lourenço, 2011*
L. kamshetensis Tikader & Bastawade, 1983
L. kharpadi Bastawade, 1986
L.
krali Kovarík, 1995
L. laevifrons
Pocock, 1897
L. Lourençoi Kovarík,
1997
L.
marmoreus (C.L. Koch, 1844)
L.
mjobergi Kraepelin, 1916
L. mucronatus
(Fabricius. 1798)
L. nigristernis (Pocock,
1899)
L. obsti Kraepelin, 1913
L.
perfidus (Keyserling, 1885)
L. rackae
Kovarík, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. shoplandi (Oates, 1888)
L. srilankensis Lorenco, 1997
L. tricarinatus (Simon, 1884)
L. variatus (Thorell, 1876) |
|
Lychasioides
Vachon, 1974
L. amieti Vachon, 1974 |
Mauritanobuthus Qi & Lourenço, 2007*
M. geniezi Qi &Lourenço, 2007*
|
Mesobuthus Vachon,
1950
M. agnetis (Werner, 1936)
M. bolensis Sun, Zhu & Lourenço, 2010*
M. caucasicus (Nordmann, 1840)
M. cyprius Gantenbein & Kropf, 2000*
M. eupeus (C.L. Koch, 1839)
M. extremus (Werner, 1936)
M. gibbosus (Brulli, 1832)
M. karshius Sun & Sun, 2011*
M. longichelus Sun & Zhu, 2010*
M. macmahoni (Pocock, 1900)
M. martensii (Karsch, 1879)
M. nigrocinctus (Ehrenberg, 1828)*
M. phillipsi (Pocock, 1889)*
M. vesiculatus (Pocock,
1899)
|
Mesotityus
Gonzalez-Sponga, 1981
M. vondangeli
González-Sponga, 1981 |
Microananteris
Lourenço, 2003*
M. minor Lourenço, 2003* |
Microbuthus
Kraepelin, 1898
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. gardneri Lowe, 2010*
M. kristensenorum Lowe, 2010*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
M. pusillus Kraepelin, 1898 |
Microcharmus Lourenço, 1995
M. bemaraha Lourenço, Goodman & Fisher, 2006*
M. cloudsleythompsoni Lourenço, 1995
M. confluenciatus Lourenço, Goodman & Fisher, 2006*
M. duhemi Lourenço, Goodman & Fisher, 2006*
M. fisheri Lourenço, 1998
M. hauseri Lourenço, 1996
M. jussarae Lourenço, 1996*
M. maculatus Lourenço, Goodman & Fisher, 2006*
M. madagascariensis Lourenço, 1999*
M. pauliani (Lourenço, 2004)*
M. sabineae Lourenço, 1996
M. variegatus Lourenço, Goodman & Fisher, 2006*
M. violaceous Lourenço, Goodman & Fisher, 2006*
|
Microtityus
Kjellesvig-Waering, 1966
M. angelaerrosae Gonzáles-Sponga, 2001*
M. barahona Armas & Teruel, 2012*
M. biordi González-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. capayaensis González-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae González-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. farleyi Teruel, 2000*
M. flavescens Teruel, 2001*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M. jaumei Armas, 1974
M. joseantonioi González-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis González-Sponga, 2001*
M. lourencoi Armas & Teruel, 2012*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. prendini Armas & Teruel, 2012*
M. pusillus Teruel & Kovarik, 2012*
M. reini Armas & Teruel, 2012*
M. rickyi Kjellesvig-Waering, 1966
M. sevciki González-Sponga, 2001*
M. solegladi Armas & Teruel, 2012*
M. starri Lourenço & Huber, 1999*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. virginiae Armas, 1999*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus González-Sponga, 2001*
|
Neobuthus Hirst,
1911
N. awashensis Kovarik & Lowe, 2012*
N. cloudsleythompsoni Lourenço, 2001*
N. berberensis (Hirst, 1911)
N. ferrugineus (Kraepelin, 1898)*
N. sudanensis Lourenço, 2005*
|
Neogrosphus
Lourenço, 1995
N. blanci Lourenço,
1996
N. griveaudi (Vachon, 1969) |
Neoprotobuthus Lourenço, 2000*
N. intermedius Lourenço, 2000* |
Odontobuthus Vachon, 1950
O. bidentatus (Lourenço & Pezier, 2002)*
O. brevidigitus Lowe, 2010*
O. doriae (Thorell, 1876)
O. odonturus (Pocock, 1897)
|
Odonturus Karsch,
1879
O.
dentatus Karsch, 1879 |
Orthochiroides
Kovarík, 1998
O. insularis (Pocock, 1889)*
O. socotrensis Kovarík, 2004*
O. vachoni Kovarík, 1998
|
Orthochirus
Karsch, 1891
O. afghanus Kovarik, 2004*
O. aristidis (Simon, 1882)
O. atarensis Lourenço & Leguin, 2011*
O. bastawadei Zambre, Mirza, Sanap, Upadhye & Javed, 2011*
O. bicolor (Pocock, 1897)
O. blandini (Lourenço & Vachon, 1997)*
O. cloudsleythompsoni Lourenço & Leguin, 2011*
O. danielleae (Lourenço & Vachon, 1997)*
O. erardi (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fuscipes (Pocock, 1900)
O. glabrifrons(Kraepelin, 1903)*
O. goyffoni (Lourenço & Vachon, 1995)*
O. gromovi Kovarik, 2004*
O. gruberi Kovarik & Fet, 2006*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. kaspareki (Lourenço & Huber, 2000)*
O. kinzelbachi (Lourenço & Huber, 2000)*
O. krishnai Tikader & Bastawade, 1983
O. maroccanus Lourenço & Leguin, 2011*
O. minor Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. monodi (Lourenço & Vachon, 1997)*
O. pallidus (Pocock, 1897)
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. stockwelli (Lourenço & Vachon, 1995)*
O. tassili Lourenço & Leguin, 2011*
O. tibesti Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. varius Kovarik, 2004*
O. zagrosensis Kovarik, 2004*
|
Pantobuthus
Lourenço & Duhem, 2009*
P. complicatus Lourenço & Duhem, 2009*
|
Parabuthus Pocock,
1890
P. brevimanus (Thorell, 1876)
P. calvus Purcell, 1898
P. capensis (Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral, 1980
P. eritreaensis Kovarik, 2003*
P. glabrimanus Prendini & Esposito, 2010*
P. gracilis Lamoral, 1979
P. granimanus Pocock, 1895
P.
granulatus (Ehrenberg, 1831)
P.
heterurus Pocock, 1899
P. hunteri
Pocock, 1895
P. kalaharicus Lamoral,
1977
P. kraepelini Werner, 1902
P.
kuanyamarum Monrad, 1937
P. laevifrons
(Simon, 1888)
P.
liosoma (Ehrenberg, 1828)
P. maximus Werner, 1913
P. mossambicensis (Peters, 1861)
P. muelleri Prendini, 2000*
P. namibensis Lamoral, 1979
P. nanus Lamoral, 1979
P. pallidus Pocock, 1895
P. planicauda (Pocock, 1889)
P. raudus (Simon, 1888)
P. schlechteri Purcell, 1899
P. setiventer Prendini & Esposito, 2010*
P. stridulus Hewitt, 1913
P.
transvaalicus Purcell, 1899
P.
villosus (Peters, 1862)
|
Pectinibuthus Fet,
1984
P. birulai Fet, 1984 |
Physoctonus
Mello-Leitao, 1934*
P. debilis (C. L. Koch, 1840)*
|
Picobuthus Lowe, 2010*
P. dundoni Lowe, 2010*
P. wahibaensis Lowe, 2010*
|
Plesiobuthus
Pocock, 1900
P. paradoxus Pocock, 1900
|
Polisius Fet,
Capes & Sissom, 2001*
P. persicus Fet,
Capes & Sissom, 2001* |
Psammobuthus
Birula, 1911
P. zarudnyi Birula, 1911
|
Pseudolissothus
Lourenço, 2001*
P. pussilus Lourenço, 2001*
|
Pseudolychas
Kraepelin, 1911
P. ochraceus (Hirst,
1911)
P. pegleri (Purcell, 1901)
P.
transvaalicus Lawrence, 1961
|
|
Pseudouroplectes
Lourenço, 1995
P. betschi Lourenço, 1995
P. lalyae Lourenço & Ythier, 2010*
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999* |
Razianus Farzanpay, 1987
R. xinjianganus Lourenco, Sun & Zhu, 2010*
R. zarudnyi (Birula, 1903)
|
Rhopalurus Thorell, 1876
R. abudi Armas & Marcano Fondeur, 1987
R. acromelas Lutz & Mello, 1922
R. agamemmon (C. L. Koch, 1839)
R. amazonicus Lourenço, 1986
R. aridicola Teruel & Armas, 2012*
R. bonettii Armas 1999*
R. crassicauda Caporiacco, 1947*
R. garridoi Armas, 1974
R. guanambiensis Lenarducci, Pinto-da-Rocha & Lucas, 2005*
R. gibarae Teruel, 2006*
R. granulimanus Teruel, 2006*
R. junceus (Herbst, 1800)
R. lacrau Lourenço & Pinto-da-Rocha, 1997
R. laticauda Thorell, 1876
R. melloleitaoi Teruel & Armas, 2006*
R. pintoi Mello-Leităo, 1932*
R. princeps (Karsch, 1879)
R. rochai Borelli, 1910
R. virkkii Santiago-Blay, 2009*
|
Riftobuthus
Lourenço, Duhem & Cloudsley-Thompson 2010*
R. inexpectatus Lourenço, Duhem & Cloudsley-Thompson 2010*
|
Sabinebuthus
Lourenço, 2001*
S. elegans Lourenço, 2001*
|
Saharobuthus
Lourenço & Duhem, 2009*
S. elegans Lourenço & Duhem, 2009*
|
Sassandiothus
Farzanpay, 1987
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900)
|
Somalibuthus
Kovarík, 1998
S. demisi Kovarík, 1998 |
Somalicharmus
Kovarík, 1998
S. withmanae Kovarík, 1998 |
Thaicharmus
Kovarík, 1995
T. lowei Kovarík, Soleglad & Fet, 2007*
T. mahunkai Kovarík, 1995
|
Tityobuthus
Pocock, 1893
T. antsingy Lourenço & Goodman, 2004*
T. baroni (Pocock, 1890)
T. betschi Lourenço, Qi & Goodman, 2008*
T. chelbergorum Lourenço, Qi & Goodman, 2008*
T. darainensis Lourenço & Goodman, 2002*
T. dastychi Lourenço, 1997
T. griswoldi Lourenço, 2000*
T. guillaumeti
Lourenço, 1995
T. ivohibe Lourenço & Goodman, 1999*
T. judsoni Lourenço,
1996
T. manonae Lourenço, 2000*
T. mccarteri Lourenço, Qi & Goodman, 2008*
T. monodi Lourenço, 2000*
T. pallidus Lourenço, 2004*
T.
parrilloi Lourenço, 1996
T. petrae
Lourenço, 1996
T. pococki Lourenço, 1995
T. rakotondravonyi Lourenço, 2003*
|
Tityopsis Armas,
1974
T. aliciae Armas & Frias, 1998*
T. inaequalis (Armas, 1974)
T.
inexpectata (Moreno, 1940) |
Tityus C.L. Koch,
1836
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. abudi Armas, 1999*
T. acananensis González-Sponga, 2009*
T. adisi Lourenço, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi González-Sponga, 2001*
T. altithronus Armas, 1999*
T. anasilviae Armas & Abud Antun, 2004*
T. androcottoides (Karsch, 1879)
T. anduzei González-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. antioquensis Lourenço& Otero Patińo, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai González-Sponga, 1985
T. argentinus Borelli,
1899
T. asthenes Pocock, 1893
T.
atriventer Pocock, 1897
T. bahiensis
(Perty, 1833)
T. bahoruco Teruel & Armas, 2006*
T. barquisimetanus
González-Sponga, 1994
T. bastosi Lourenço,
1984
T. bellulus Armas, 1999*
T.
betschi Lourenço, 1992
T. birabeni
Abalos, 1955
T. blanci Lourenço,
1994
T. blaseri Mello-Leităo, 1931
T. boconoensis González-Sponga, 1981
T.
bolivanus Kraepelin, 1895
T. brazilae
Lourenço & Eickstedt, 1984
T. breweri
González-Sponga, 1997
T. cachipalensis González-Sponga, 2002*
T. caesarbarrioi González-Sponga, 2001*
T. canopensis Lourenço & Pezier, 2002*
T. carabobensis González-Sponga, 1987
T. carinatoides Mello-Leităo, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carvalhoi Mello-Leităo, 1945
T. cerroazul Lourenço, 1986
T. championi Pocock, 1898*
T. charalaensis Mello-Leităo, 1940
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. clathratus C. L. Koch, 1844
T.
columbianus (Thorell, 1876)
T.
confluens Borelli, 1899
T. costatus
(Karsch, 1879)
T. crassimanus (Thorell,
1876)
T. cuellari Lourenço, 1994
T.
culebrensis González-Sponga, 1994
T. cylindricus (Karsch, 1879)*
T.
dasyurus Pocock, 1897
T. dedoslargos
Francke & Stockwell, 1987
T. demangei
Lourenço, 1981
T. dinizi Lourenço,
1997
T. discrepans (Karsch, 1879)
T. dorae González-Sponga, 2001*
T. duacaensis González-Sponga, 2007*
T. dulceae González-Sponga, 2006*
T. dupouyi González-Sponga, 1987
T.
ebanoverde Armas, 1999*
T. ecuadorensis
Kraepelin, 1896
T. elii (Armas & Marcano
Fondeur, 1992)*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi González-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leităo, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessôa, 1935
T. festae Borelli, 1899
T. filodendron González-Sponga, 1981
T. florezi Lourenço, 2000*
T. footei Chamberlin, 1916
T. forcipula (Gervais, 1843)
T. fuhrmanni Kraepelin, 1914
T. funestus Hirst, 1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. grahami Lourenço, 2012*
T. guariconesis Gonzalez-Sponga, 2004*
T. horacioi Lourenço& Leguin, 2011*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus Mello-Leităo, 1934
T. insignis (Pocock, 1889)
T. intermedius Borelli, 1899
T. irapaensis González-Sponga, 2002*
T. isabelceciliae González-Sponga, D'Suze & Sevcik, 2001*
T. ivani González-Sponga, 2008*
T. ivicnancor González-Sponga,
1997
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai González-Sponga, 2007*
T. kuryi Lourenço, 1997
T. lancinii González-Sponga, 1972
T. lokiae Lourenço, 2005*
T. longidigitus González-Sponga, 2008*
T. Lourençoi Flórez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis González-Sponga, 2007*
T. mana Lourenço, 2012*
T. manakai González-Sponga, 2004*
T. maniapurensis González-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. martinpaechi Lourenço, 2001*
T. matthieseni Lourenço & Pinto-da-Rocha, 2000
T. mattogrossensis Borelli,
1901
T. maturinensis González-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis González-Sponga, 1974
T. mongei Lourenço, 1996
T. mucusunamensis González-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leităo, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leităo, 1940
T. neoespartanus González-Sponga, 1996
T. nororientalis González-Sponga, 1996
T. obispoi González-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus González-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis González-Sponga, 2007*
T. paraguayensis Kraepelin, 1895
T.
parvulus Kraepelin, 1914
T. paulistorum Lourenço & Qi, 2006*
T.
perijanensis González-Sponga, 1994
T.
pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri
González-Sponga, 1981
T. pococki Hirst,
1907
T. portoplatensis Armas & Marcano
Fondeur, 1992
T. potameis Lourenço & Leăo Giupponi, 2004*
T. prancei Lourenço,
2000*
T. proseni Abalos, 1954
T.
pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis González-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus (Armas, 1982)*
T. ramirezi Esquivel de Verde, 1969 [nomen dubium]
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis González-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi González-Sponga, 1996
T. romeroi González-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rufofuscus Pocock, 1897
T. rugosus Schenkel, 1932
T. rusmelyae González-Sponga, D'Suze & Sevcik, 2001*
T. sabinae Lourenço, 1994
T. sanarensis González-Sponga, 1997
T. sanfernandoi González-Sponga, 2008*
T. sarisarinamensis González-Sponga, 2002*
T. sastrei Lourenço & Flórez, 1990
T. septentrionalis Armas & Abud Antun, 2004*
T. serrulatus Lutz & Mello, 1922
T. shiriana González-Sponga, 1991
T. silvestris Pocock, 1897
T. simonsi Pocock, 1900
T. smithii Pocock, 1893
T. soratensis Kraepelin, 1912
T. stigmurus (Thorell, 1876)
T. strandi Werner, 1939
T. surimeridensis González-Sponga, 2002*
T. surorientalis González-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi González-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. thelyacanthus Mello-Leităo, 1933
T. timendus Pocock, 1898
T. trinitatis Pocock, 1897
T. trivittatus Kraepelin, 1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leităo, 1931
T. unus Lourenço & Pinto-da-Rocha, 2000
T. uquirensis González-Sponga, 2001*
T. urachichensis González-Sponga, 2007*
T. urbinai
Scorza, 1952
T. uruguayensis Borelli,
1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis González-Sponga, 1981
T. ventuarensis González-Sponga, 2009*
T. walli González-Sponga & Wall Gonzalez, 2007*
T. wayuu Rojas-Runjaic & Armas, 2007*
T. yerenai González-Sponga, 2009*
T. ythieri Lourenço, 2007*
T. zulianus González-Sponga, 1981
|
|
Troglorhopalurus Lourenço, Baptista & Giupponi,
2004*
T. translucidus Lourenço, Baptista & Giupponi,
2004*
| Troglotityobuthus Lourenço,
2000*
T. gracilis (Fage, 1946)
|
Uroplectes Peters,
1861
U. fischeri (Karsch, 1879)
U.
andreae Pocock, 1889
U. carinatus
(Pocock, 1890)
U. chubbi Hirst,
1911
U. fischeri (Karsch, 1879)
U.
flavoviridis Peters, 1861
U. formosus
Pocock, 1890
U. gracilior Hewitt,
1914
U. insignis Pocock, 1890
U.
lineatus (C. L. Koch, 1844)
U.
longimanus Werner, 1936
U. marlothi
Purcell, 1901
U. ngangelarum Monard,
1930
U. occidentalis Simon, 1876
U.
olivaceus Pocock, 1896
U.
otjimbinguensis (Karsch, 1879)
U.
pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner,
1913
U. pilosus (Thorell, 1876)
U.
planimanus (Karsch, 1879)
U.
schlechteri Purcell, 1901
U. schubotzi
Kraepelin, 1929
U. silvestrii Borelli,
1913
U. teretipes Lawrence, 1966
U.
triangulifer (Thorell, 1876)
U.
tumidimanus Lamoral, 1979
U. variegatus
(C. L. Koch, 1844)
U. vittatus (Thorell,
1876)
U. xanthogrammus Pocock,
1897
| |
Uroplectoides
Lourenço, 1998
U. abyssinicus Lourenço,
1998
U. emiliae (Werner, 1916) |
Vachoniolus Levy, Amitai & Shulov, 1973
V. batinahensis Lowe, 2010*
V. gallagheri Lowe, 2010*
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
|
Vachonus Tikader & Bastawade, 1983
V. asiyaae Amir & Kamaluddin, 2009*
V. iqbali Amir & Kamaluddin, 2009*
V. rajasthanicus Tikader & Bastawade, 1983
|
Zabius Thorell, 1893
Z. birabeni Mello-Laeităo, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*
|
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