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Hemiscorpiidae
Pocock, 1893 * |
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This family has previously been known
as Ischnuridae Simon, 1879, but in ICZN. 2003, Opinion 2037 (cases 3120 and 3120a), the name has been changed to
Liochelidae Fet & Bechly, 2001. The reason for this change is that the old name was in conflict with a Dragonfly
(Odonata) subfamily name (Ischnurinae). More info in Bulletin of Zoological Nomenclature, 60(2): 159-161 (June 2003). In 2005, Soleglad, Fet & Kovarik changed the name to Hemiscorpiidae, and included the genus Heteroscorpion (from the abolished family Urodacidae) in addition to all genera and species previously included in Liochelidae.
In a major revision of higher scorpion systematics, Soleglad & Fet (2003) included the genera Habibella and Hemiscorpius (from the now abolished family Hemiscorpiidae) into Liochelidae (now Hemiscorpiidae).
The genus Habibella was synonymized with Hemiscorpius by Monod & Lourenço in late 2005.
The genus Liocheles was revised by Monod in late 2011.
This family consists of 12 genera and 94 species [Updated 11.12.12], and
inhabits all continents except North America. Hemiscorpiid
scorpions are found in tropical and partly in subtropical
habitats. Heteroscorpion is endemic for Madagascar.
Some of the scorpions in this family are quite common in
captivity. In particular, the large and fascinating members of
the genus Hadogenes are popular, but these are
difficult to get due to limited numbers in the pet trade. Also
some species of Iomachus and Opistacanthus
have been reported in captivity.
The scorpions in this family are not medically important (except Hemiscorpius),
and some of the species have very high LD50 values. (High LD50
value means less venomous; e.g., Hadogenes
troglodytes has a reported LD50 of 2,000 mg/kg (LD50 =
Lethal Dose in 50% of samples, milligram venom/kilogram rat or
mouse weight: Leiurus has an LD50 well below 1.00
mg/kg, for comparison)). Many of the members of this family
have very slender caudas (tails, metasomas).
NB! Hemiscorpius lepturus is a very dangerous species
of serious medical significance (see species link for more
information). No data have been available for other members of
this genus, but they should be handled with extreme
care as long as their venom status is unknown.
A characteristic of many members of family, especially,
Hadogenes, is that the scorpions are very flat (looks
like somebody has stomped on them). This is an adaption to a
life in cracks and crevices. They resemble the scorpions in
the family Scorpionidae, in which they were previously
included. Some members of this family are very slow-growing,
long lived, and can attain very long lengths (+ 20 cm).
Iomachus politus photo (left) by Jan
Ove Rein (C). Hadogenes troglodytes photo (right)
by Jan Ove Rein (C).
SPECIES
FILES: This list of genera and species is
based on Fet et al.(2000). Subspecies is not included in the list.
I try to update the list as additions and changes are published. * denotes changes after Fet et al. (2000). I will be grateful
for information about new development in the taxonomy of this family.
For information about synonyms and bibliographies, see Fet et al. (2000).
See also Soleglad & Fet (2003) for more information. Fulltext freely available HERE (issue 11).
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Cheloctonus Pocock, 1892
C. anthracinus Pocock, 1899
C. crassimanus (Pocock, 1896)
C. glaber Kraepelin, 1896
C. intermedius Hewitt, 1912
C. jonesii Pocock, 1892
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Chiromachetes Pocock, 1899
C. fergusoni Pocock, 1899
C. tirupati Lourenço, 1997
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Chiromachus Pocock, 1893
C. ochropus (C. L. Koch, 1837)
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Hadogenes Kraepelin, 1894
H. austroafricanus Penther, 1900
H. bicolor Purcell, 1899
H. gracilis Hewitt, 1909
H. granulatus Purcell, 1901
H. gunnigi Purcell, 1899
H. hahni (Peters, 1862)* (= H. taeniurus)
H. lawrencei Newlands, 1972
H. longimanus Prendini, 2001*
H. minor Purcell, 1899
H. newlandsi Prendini, 2001*
H. paucidens Pocock, 1896
H. phyllodes Thorell, 1876
H. polytrichobothrius Prendini, 2006*
H. southpansbergensis Prendini, 2006*
H. tityrus (Simon, 1888)
H. trichiurus (Gervais, 1843)
H. troglodytes (Peters, 1861)
H. zuluanus Lawrence, 1937
H. zumpti Newlands & Cantrell, 1985
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Hemiscorpius Peters, 1861
H. acanthocercus Monod & Lourenço, 2005*
H. arabicus (Pocock, 1899)
H. egyptiensis Lourenço, 2011*
H. enischnochela Monod & Lourenço, 2005*
H. falcifer Lowe, 2010*
H. flagelliraptor Loewe, 2010*
H. gaillardi (Vachon, 1974)*
H. kashkayi Karatas & Gharkheloo, 2013*
H. lepturus Peters, 1861
H. maindroni (Kraepelin, 1900)
H. novaki Kovarik & Mazuch, 2011*
H. persicus Birula, 1903
H. socotranus Pocock, 1899
H. somalicus Lourenço, 2011*
H. tellinii Borelli, 1904
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Heteroscorpion Birula, 1903
H. goodmani Lourenço, 1996
H. kaii Lourenço & Goodman, 2009*
H. kraepelini Lourenço & Goodman, 2006*
H. magnus Lourenço & Goodman, 2002*
H. opisthacanthoides (Kraepelin, 1896)
H. raselimananai Lourenço & Goodman, 2004*
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Hormiops Fage, 1933*
H. davidovi Fage, 1933*
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Iomachus Pocock, 1893
I. laeviceps (Pocock, 1890)
I. nitidus Pocock, 1896
I. politus Pocock, 1896
I. punctulatus Pocock, 1897
I. surgani (Bastawade, 1986
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Liocheles Sundevall, 1833
L. australasiae (Fabricius, 1775)
L. boholiensis (Kraepelin, 1914)*
L. extensus Locket, 1997
L. karschii (Keyserling, 1885)
L. litodactylus Monod & Volschenk, 2004*
L. longimanus (Werner, 1939)*
L. neocaledonicus (Simon, 1877)
L. nigripes (Pocock, 1897)
L. penta Francke & Lourenço, 1991
L. polisorum Volschenk, Locket & Harvey, 2001*
L. waigiensis (Gervais 1843)
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Opisthacanthus Peters, 1861
O. africanus Simon, 1876
O. asper (Peters, 1861)
O. autanensis González-Sponga, 2004*
O. basutus Lawrence, 1955
O. borboremai Lourenço & Fe, 2003*
O. brevicauda Rojas-Runjaic, Borges & Armas, 2008*
O. capensis Thorell, 1876
O. cayaporum Vellard, 1932
O. darainensis Lourenço & Goodman, 2006*
O. diremptus (Karsch, 1879)
O. elatus (Pocock, 1893)
O. laevipes Vellard, 1932
O. lamorali Lourenço, 1981
O. lecomtei (Lucas, 1858)
O. lepturus (Beauvois, 1805)
O. lucienneae Lourenço & Goodman, 2006*
O. maculatus Lourenço & Goodman, 2006*
O. madagascariensis Kraepelin, 1894
O. milloti Lourenço & Goodman, 2008*
O. pauliani Lourenço & Goodman, 2008*
O. piceus Lourenço & Goodman, 2006*
O. piscatorius Lawrence, 1955
O. rugiceps Vellard, 1932
O. rurgulosus Pocock, 1896
O. valerioi Lourenço, 1980
O. validus Thorell, 1876
O. weyrauchi Mello-Leitão, 1948
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Paleocheloctonus Lourenço, 1996
P. pauliani Lourenço, 1996
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Tibetiomachus Lourenço & Qi, 2006*
T. himalayensis Lourenço & Qi, 2006*
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